<document id="CAB7C528CBA6B4122D0901A9DEB94C0D" ID-CLB-Dataset="37646" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE48FFD67DB5FE551392DE75" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes" docType="key" docVersion="11" lastPageNumber="57" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="57" updateTime="1732884161890" updateUser="ExternalLinkService">
<mods:mods id="DE078A74EE65DA48ECA3CA3EB352464C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A35ECE5F57961B991105F1A8C4EC0417">
<mods:title id="28215FA8D120F1197716B53CAD58E0C9">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="13ABAE246904B0655457EC3AED0C5C0C" type="personal">
<mods:role id="8F407B45B8EB7A4BBD81ED675E918132">
<mods:roleTerm id="708856E14CE54BDF923C3E22D0638764">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7673DD364724C3DC904B0E05A6B7479D">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="D77E896A7FE0D66673E4048C5BE6119D" type="personal">
<mods:role id="77B0561969A21971560E705013709D4D">
<mods:roleTerm id="DBD3539AF3B18C69ED4879BF38D85245">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56B1D2157D5FBDF2EFC2957A50D3E360">Spangler, Paul J.</mods:namePart>
</mods:name>
<mods:name id="FC63BB9DAE5FFAC099F314B0B87A4897" type="personal">
<mods:role id="9EFF40D3B17BC0EC475F5C1A0B86758F">
<mods:roleTerm id="772308DBE0410B4D1EC0FC834FCDC792">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2236AB97678828AD0DFAA1F4D16D09E2">Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="4F5F5B80CDDEBAE3B58A90E7D0741B96">text</mods:typeOfResource>
<mods:relatedItem id="CB7D4A818E3A2A326E4C332CBDEDB675" type="host">
<mods:titleInfo id="9BAD83B62A9B516E47D20856CB999285">
<mods:title id="7D2CA9F4335867CDB8B027DCFEC42636">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="708E4975C146355DBD5A6544CBF68F6A">
<mods:date id="EDAE3118CA136F924D9A110C7E78C5B4">2010</mods:date>
<mods:detail id="FCC796008765B35663AE432253E712A2" type="volume">
<mods:number id="B8939A8FE2A2F2904B7F96114151F0E3">2347</mods:number>
</mods:detail>
<mods:extent id="AC6E3003786C532693A93C52CC92DF8C" unit="page">
<mods:start id="18D3303B50406DC2C52A634B41F16AF0">37</mods:start>
<mods:end id="67ABFB54D5B99300F9EBE9F8D32CA8B4">58</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="014EDB3AFA6F3DC985928B4CDA7CA914">journal article</mods:classification>
<mods:identifier id="422BED58A86921F3273FBEB39142EB8F" type="CLB-Dataset">37646</mods:identifier>
<mods:identifier id="887458F9D69E5671A61CBB600751B96A" type="DOI">10.5281/zenodo.193406</mods:identifier>
<mods:identifier id="30472710A203F2B14DF93D340D2464B4" type="GBIF-Dataset">5e8043a5-e685-4a21-b696-0f7a706ab223</mods:identifier>
<mods:identifier id="DC2C2673C71716EC7C4DE47B105DCECD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="96E5E5C5B8F3BF8C514A4BC29F604594" type="Zenodo-Dep">193406</mods:identifier>
</mods:mods>
<treatment id="11578788FE48FFD67DB5FE551392DE75" ID-DOI="http://doi.org/10.5281/zenodo.6198400" ID-GBIF-Taxon="124955822" ID-Zenodo-Dep="6198400" LSID="urn:lsid:plazi:treatment:11578788FE48FFD67DB5FE551392DE75" httpUri="http://treatment.plazi.org/id/11578788FE48FFD67DB5FE551392DE75" lastPageNumber="57" pageId="20" pageNumber="57" scope_family="Dytiscidae" scope_order="Coleoptera">
<subSubSection id="D1E46515FE48FFD67DB5FE5517FCD992" box="[151,498,422,448]" pageId="20" pageNumber="57" type="nomenclature">
<paragraph id="9941369EFE48FFD67DB5FE5517FCD992" blockId="20.[151,498,422,448]" box="[151,498,422,448]" pageId="20" pageNumber="57">
<heading id="C20981F2FE48FFD67DB5FE5517FCD992" bold="true" box="[151,498,422,448]" fontSize="11" level="1" pageId="20" pageNumber="57" reason="1">
<emphasis id="AB8AEA8CFE48FFD67DB5FE5517FCD992" bold="true" box="[151,498,422,448]" pageId="20" pageNumber="57">
Key to species of 
<taxonomicName id="5EFE4D1DFE48FFD67C4FFE5517FCD992" ID-CoL="59RJ" box="[365,498,422,448]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="AB8AEA8CFE48FFD67C4FFE5517FCD992" bold="true" box="[365,498,422,448]" italics="true" pageId="20" pageNumber="57">Laccodytes</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE48FFD67DB5FE031392DE75" pageId="20" pageNumber="57" type="key">
<keyStep id="220A2E3BFE48FFD67DB5FE031395DA35" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FE031395DA75" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FE031395DA75" pageId="20" pageNumber="57">1. Posterior angles of pronotum produced backwards, acute; prosternal process with very long, needle-shaped tip, reaching caudally beyond mesocoxae for about a third of its length ........................................................................... 2</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FDC31395DA35" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FDC31395DA35" pageId="20" pageNumber="57">- Posterior angles of pronotum rounded; prosternal process with much shorter thorn-shaped tip, not or slightly extended caudally beyond mesocoxae .......................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FD831395DB75" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FD831392DA95" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FD831392DA95" pageId="20" pageNumber="57">
2. Dark line along suture complete, reaching the scutellum, fused laterally with humeral dark spots; subbasal band reaching the sides of elytra; dark patterns brown. Median lobe of aedeagus slender and more simple in shape........... 
<emphasis id="AB8AEA8CFE48FFD67DECFD431392DA95" box="[206,1436,688,711]" italics="true" pageId="20" pageNumber="57">
..................................................................................................................................................................... 
<taxonomicName id="5EFE4D1DFE48FFD6783DFD431392DA95" ID-CoL="3RLZF" box="[1311,1436,688,711]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="apalodes">L. apalodes</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FD231395DB75" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FD231395DB75" pageId="20" pageNumber="57">
- Dark line along suture not reaching the scutellum, ending in a lobed shape; humeral spots isolated; subbasal band not reaching the sides of elytra; dark patterns black. Median lobe of aedeagus larger and robust, more complex in shape 
<emphasis id="AB8AEA8CFE48FFD67C28FCE31395DB75" box="[266,1435,784,807]" italics="true" pageId="20" pageNumber="57">
............................................................................................................................................................ 
<taxonomicName id="5EFE4D1DFE48FFD6783DFCE31395DB75" ID-CoL="3RLZN" box="[1311,1435,784,807]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="rondonia">L. rondonia</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FCC31395DBF5" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FCC31392DB35" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FCC31392DB35" pageId="20" pageNumber="57">
3. MR on metacoxae composed of elongate, fine longitudinal cells, visible at high magnification, punctuation not visible; elytra regularly tapering posteriourly, rounded apically 
<emphasis id="AB8AEA8CFE48FFD67E62FCA31392DB35" box="[832,1436,848,871]" italics="true" pageId="20" pageNumber="57">
................................................................... 
<taxonomicName id="5EFE4D1DFE48FFD67821FCA31392DB35" ID-CoL="3RLZD" box="[1283,1436,848,871]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="americanus">L. americanus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FC831395DBF5" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FC831395DBF5" pageId="20" pageNumber="57">- MR on metacoxae composed of rounded meshes, together with a more or less visible punctuation of fine dots; elytra rounded or truncate apically......................................................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FC431395DBB5" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FC431392DB95" blockId="20.[151,1436,496,1575]" box="[151,1436,944,967]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FC431392DB95" box="[151,1436,944,967]" pageId="20" pageNumber="57">
4. Size larger (&gt; 
<quantity id="5E069B7BFE48FFD67C42FC4317ACDB95" box="[352,418,944,967]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="57" unit="mm" value="2.0">2 mm</quantity>
); Median lobe of aedeagus large and robust ........................................................ 
<taxonomicName id="5EFE4D1DFE48FFD679D7FC431392DB95" ID-CoL="3RLZL" box="[1269,1436,944,967]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="phalacroides">
<emphasis id="AB8AEA8CFE48FFD679D7FC431392DB95" box="[1269,1436,944,967]" italics="true" pageId="20" pageNumber="57">L. phalacroides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FC231395DBB5" blockId="20.[151,1436,496,1575]" box="[151,1435,976,999]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FC231395DBB5" box="[151,1435,976,999]" pageId="20" pageNumber="57">
- Size smaller (&lt;
<quantity id="5E069B7BFE48FFD67C53FC2317BDDBB5" box="[369,435,976,999]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="57" unit="mm" value="2.0">2 mm</quantity>
); Median lobe of aedeagus small and slender ........................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FC031395DC35" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FC03139BDC75" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FC03139BDC75" pageId="20" pageNumber="57">
5. Apex of each elytron obliquely truncate, almost concave in outline; elytral patterns sharply contrasting: yellow subbasal and apical bands deeply lobate, giving to the dark markings a strongly irregular appearance 
<emphasis id="AB8AEA8CFE48FFD679F5FBE3139BDC75" box="[1239,1429,1040,1063]" italics="true" pageId="20" pageNumber="57">
...... 
<taxonomicName id="5EFE4D1DFE48FFD67826FBE3139BDC75" ID-CoL="3RLZP" box="[1284,1429,1040,1063]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="takutuanus">L. takutuanus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FBC31395DC35" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FBC31395DC35" pageId="20" pageNumber="57">- Apex of each elytron not truncate, otherwise not obliquely; dark patterns less contrasting or, if so, with markings not lobate............................................................................................................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FB831395DC95" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FB831392DCD5" blockId="20.[151,1436,496,1575]" box="[151,1436,1136,1159]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FB831392DCD5" box="[151,1436,1136,1159]" pageId="20" pageNumber="57">
6. Outer metatibial spur truncate at tip. Elytra black, with three small orange-reddish circular spots 
<emphasis id="AB8AEA8CFE48FFD679E8FB831392DCD5" box="[1226,1436,1136,1159]" italics="true" pageId="20" pageNumber="57">
........... 
<taxonomicName id="5EFE4D1DFE48FFD67834FB831392DCD5" ID-CoL="3RLZK" box="[1302,1436,1136,1159]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="olibroides">L. olibroides</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FB631395DC95" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FB631395DC95" pageId="20" pageNumber="57">- Outer metatibial spur acuminate. Dark patterns of elytra brown, with a transverse sub-basal yellow band, often poorly distinct ............................................................................................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FB231395DD75" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FB231392DCB5" blockId="20.[151,1436,496,1575]" box="[151,1436,1232,1255]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FB231392DCB5" box="[151,1436,1232,1255]" pageId="20" pageNumber="57">
7. Outer metatibial spur flat. Elytra brown with poorly distinct patterns 
<emphasis id="AB8AEA8CFE48FFD67E86FB231392DCB5" box="[932,1436,1232,1255]" italics="true" pageId="20" pageNumber="57">
......................................................... 
<taxonomicName id="5EFE4D1DFE48FFD67806FB231392DCB5" ID-CoL="3RLZH" box="[1316,1436,1232,1255]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="neblinae">L. neblinae</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FB031395DD75" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FB031395DD75" pageId="20" pageNumber="57">- Both metatibial spurs conical. Elytra with more or less distinct pale patterns, otherwise colouration dark also on head and pronotum........................................................................................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FAC31392DDD5" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FAC3139ADD15" blockId="20.[151,1436,496,1575]" box="[151,1428,1328,1351]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FAC3139ADD15" box="[151,1428,1328,1351]" pageId="20" pageNumber="57">8. Head and pronotum completely or mostly dark. Median lobe of aedeagus very elongate and thin in lateral view ... 9</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FAA31392DDD5" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FAA31392DDD5" pageId="20" pageNumber="57">
- Head uniformly orange and pronotum of the same colour, at most with a dark basal line. Median lobe of aedeagus more robust in lateral view 
<emphasis id="AB8AEA8CFE48FFD67CC4FA831392DDD5" box="[486,1436,1392,1415]" italics="true" pageId="20" pageNumber="57">
....................................................................................................................... 
<taxonomicName id="5EFE4D1DFE48FFD67827FA831392DDD5" ID-CoL="3RLZG" box="[1285,1436,1392,1415]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="bassignanii">L. bassignanii</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="220A2E3BFE48FFD67DB5FA631392DE75" pageId="20" pageNumber="57">
<paragraph id="9941369EFE48FFD67DB5FA631398DDB5" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FA631398DDB5" pageId="20" pageNumber="57">
9. Dorsal colouration almost uniformly pitch-brown, with very indistinct paler patterns on elytra. Median lobe, in lateral view, visibly curved downward on apical third Male with last stenite of abdomen deeply emarginated on hind margin ..................................................................................................................................................... 
<taxonomicName id="5EFE4D1DFE48FFD67823FA231398DDB5" ID-CoL="6NRKR" box="[1281,1430,1488,1511]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="androginus">
<emphasis id="AB8AEA8CFE48FFD67823FA231398DDB5" box="[1281,1430,1488,1511]" italics="true" pageId="20" pageNumber="57">L. androginus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9941369EFE48FFD67DB5FA031392DE75" blockId="20.[151,1436,496,1575]" pageId="20" pageNumber="57">
<keyLead id="220F95ABFE48FFD67DB5FA031392DE75" pageId="20" pageNumber="57">
- Pronotum paler than head and elytra. Median lobe of aedeagus, in lateral view, completely straight and very thin, slightly curved on apical fifth. Male with last sternite not emarginated on hind margin 
<emphasis id="AB8AEA8CFE48FFD6797DF9E31392DE75" box="[1119,1436,1552,1575]" italics="true" pageId="20" pageNumber="57">
......................... 
<taxonomicName id="5EFE4D1DFE48FFD67829F9E31392DE75" ID-CoL="3RLZJ" box="[1291,1436,1552,1575]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="57" phylum="Arthropoda" rank="species" species="obscuratus">L. obscuratus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>