<document id="77AB7A79F03D9F4E598BD1FDC2822E8F" ID-CLB-Dataset="37307" ID-DOI="10.5281/zenodo.247206" ID-GBIF-Dataset="cd615cd7-ecb8-4cc3-b074-5ddfab94649a" ID-Zenodo-Dep="247206" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484571130511" checkinUser="plazi" docAuthor="Ivan Löbl &amp; Ryo Ogawa" docDate="2016" docId="0C70F70CD738FF9AFF7BA81DDC2F40C7" docLanguage="en" docName="LinzerbiolBeitr.48.2.1339-1492.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Scaphisoma pandanum Löbl &amp; Ogawa, 2016, nov.sp." docType="treatment" docVersion="13" lastPageNumber="1395" masterDocId="F0498F74D70FFFA2FFC9AB07DD0F4264" masterDocTitle="On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH" masterLastPageNumber="1492" masterPageNumber="1339" pageNumber="1394" updateTime="1732575367156" updateUser="ExternalLinkService">
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<mods:title id="D54E766A1FB931D4409F0BC2B001AC3F">On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH</mods:title>
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<mods:name id="DB7B36AE1A9280F9A7B217E92598A61C" type="personal">
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<mods:namePart id="DFB7149A324299C5D7781E22FA2B4960">Ivan Löbl</mods:namePart>
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<mods:namePart id="3F5E34D454820EED71B7CF137F160C00">Ryo Ogawa</mods:namePart>
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<mods:date id="3CBAC80C5B124A35E9DAA69C739DD8B6">2016</mods:date>
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<mods:identifier id="E02CE078DB1303A360CCE151B4B91B9D" type="DOI">10.5281/zenodo.247206</mods:identifier>
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<treatment id="0C70F70CD738FF9AFF7BA81DDC2F40C7" ID-DOI="http://doi.org/10.5281/zenodo.4390790" ID-GBIF-Taxon="127598868" ID-Zenodo-Dep="4390790" LSID="urn:lsid:plazi:treatment:0C70F70CD738FF9AFF7BA81DDC2F40C7" httpUri="http://treatment.plazi.org/id/0C70F70CD738FF9AFF7BA81DDC2F40C7" lastPageId="56" lastPageNumber="1395" pageId="55" pageNumber="1394" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Scaphidiinae" scope_tribe="Scaphisomatini">
<subSubSection id="CCC31591D738FF95FF7BA81DDE204154" box="[178,815,793,816]" pageId="55" pageNumber="1394" type="nomenclature">
<paragraph id="8466461AD738FF95FF7BA81DDE204154" blockId="55.[178,1086,793,1603]" box="[178,815,793,816]" pageId="55" pageNumber="1394">
<emphasis id="B6AD9A08D738FF95FF7BA81DDE204154" bold="true" box="[178,815,793,816]" pageId="55" pageNumber="1394">
5.2.13.10. 
<taxonomicName id="43D93D99D738FF95FED7A81EDF1B414B" ID-CoL="6Y5DV" box="[286,532,793,815]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="1394" phylum="Arthropoda" rank="species" species="pandanum" status="sp. nov.">
<emphasis id="B6AD9A08D738FF95FED7A81EDF1B414B" bold="true" box="[286,532,793,815]" italics="true" pageId="55" pageNumber="1394">Scaphisoma pandanum</emphasis>
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<taxonomicNameLabel id="AD9E2773D738FF95FDD3A81DDF69414B" box="[538,614,794,815]" pageId="55" pageNumber="1394">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1CE25A9FD738FF95FDBAA81DDFCC414B" box="[627,707,794,816]" captionStart="Figs 22 - 28" captionStartId="113.[178,218,1394,1416]" captionText="Figs 22 - 28: Habitus of Scaphisoma; (22), S. mirum nov. sp.; (23), S. nabiluanum nov. sp.; (24), S. ochropenne nov. sp.; (25), S. opacum nov. sp.; (26), S. palawanum PIC; (27), S. pandanum nov. sp.; (28), S. pseudokalabitum nov. sp. Scale: 1.0 mm." pageId="55" pageNumber="1394">Figs 27</figureCitation>
, 
<figureCitation id="1CE25A9FD738FF95FD07A81DDE274154" box="[718,808,794,816]" captionStart="Figs 131 - 136" captionStartId="131.[178,218,1454,1476]" captionTargetBox="[207,1066,221,1421]" captionTargetId="figure@131.[178,1085,212,1444]" captionTargetPageId="131" captionText="Figs 131 - 136: Aedeagi of Scaphisoma in dorsal and lateral views; (131, 132), S. palawanum PIC; (133, 134), S. pandanum nov. sp.; (135, 136), S. scapulare nov. sp. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.247220" httpUri="https://zenodo.org/record/247220/files/figure.png" pageId="55" pageNumber="1394">133, 134</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CCC31591D738FF95FF7BA83DDFDD4119" pageId="55" pageNumber="1394" type="materials_examined">
<paragraph id="8466461AD738FF95FF7BA83DDFDD4119" blockId="55.[178,1086,793,1603]" pageId="55" pageNumber="1394">T y p e m a t e r i a l: Holotype 6: Philippines Occident. Mindoro North Pandan Island G. Cuccodoro 7.i.1997 10 m: sifting leaf &amp; fermenting fruits in forest (MHNG). Paratypes: with the same data as the holotype, 35 66, 63 ♀♀ (MHNG, BZLA).</paragraph>
</subSubSection>
<subSubSection id="CCC31591D738FF9AFF7BA88CDE714065" lastPageId="56" lastPageNumber="1395" pageId="55" pageNumber="1394" type="description">
<paragraph id="8466461AD738FF9AFF7BA88CDCC54369" blockId="55.[178,1086,793,1603]" lastBlockId="56.[178,1086,219,675]" lastPageId="56" lastPageNumber="1395" pageId="55" pageNumber="1394">Description: Length 1.34-1.43 mm, width 1.05-1.08 mm. Head, body and femora uniformly, fairly light reddish-brown, apical abdominal segments, tibiae, tarsi and antennae lighter, yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/7: IV 27/7: V 31/7: VI 29/8: VII 40/14: VIII 25/9: IX 40/13: X 40/14: XI 50/15. Pronotum very finely and sparsely punctate, with lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae impunctate. Exposed tip of scutellum minute. Elytra convex, with lateral margin carinae hardly exposed in dorsal view, apical margins truncate, inner apical angles not prominent, situated in level with outer angles; sutural margin not raised, sutural striae shallow, bent at base, extended along basal margins to form basal striae reaching basal mid-width, parallel posteriad except near apices, very finely punctate; adsutural areas narrow, flat, each with single puncture row, lateral striae impunctate. Elytral disc with punctation very fine, almost as that on pronotum, dense. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera about as long as intervals to mesocoxae, about twice as long as wide. Metaventrite not microsculptured. Median part of metaventrite flattened, without impressions, distinctly punctate, punctures between mesocoxae fairly sparse, posterior level of mesocoxae dense and coarse, bearing conspicuous pubescence. Lateral parts of metaventrite with extremely fine and sparse punctation; antecoxal puncture rows absent. Submesocoxal areas 0.06 mm, about as two thirds of intervals to metacoxae, submesocoxal lines convex, distinctly punctate, punctures not extended by striae. Metanepisternum flat, narrowed anteriad, inner margin rounded, suture deep. Tibiae straight. Abdomen with distinct or hardly visible punctulate microsculpture, punctation very fine and sparse, except on median area of sternite 1 bearing distinct punctures and conspicuous pubescence; submetacoxal areas 0.06- 0.07 mm, almost as half of intervals to apical margin of sternite, submetacoxal lines convex, distinctly punctate.</paragraph>
<paragraph id="8466461AD737FF9AFF7BAA19DE714065" blockId="56.[178,1086,219,675]" pageId="56" pageNumber="1395">
M a l e: Protarsomeres 1 to 3 distinctly widened. Aedeagus (
<figureCitation id="1CE25A9FD737FF9AFCF2AA1ADEC34350" box="[827,972,285,308]" captionStart="Figs 131 - 136" captionStartId="131.[178,218,1454,1476]" captionTargetBox="[207,1066,221,1421]" captionTargetId="figure@131.[178,1085,212,1444]" captionTargetPageId="131" captionText="Figs 131 - 136: Aedeagi of Scaphisoma in dorsal and lateral views; (131, 132), S. palawanum PIC; (133, 134), S. pandanum nov. sp.; (135, 136), S. scapulare nov. sp. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.247220" httpUri="https://zenodo.org/record/247220/files/figure.png" pageId="56" pageNumber="1395">Figs 132, 133</figureCitation>
) 0.53-0.71 mm long, asymmetrical, strongly sclerotized. Basal bulb fairly large. Articular processes large, not prominent. Apical process of median lobe split into two laminae, with large ostium in-between. Right lamina about as long as basal bulb, inflexed, subcylindrical in dorsal view and with acute tip, left lamina short. Parameres narrowed and with membranous inner margins in apical halves, denticular at mid-length (dorsal view). Internal sac with robust, strongly sclerotized flagellum proximally bent and hook-like, dilated and shovel-like at apex; spines and scale-like structures absent.
</paragraph>
</subSubSection>
<subSubSection id="CCC31591D737FF9AFF7BA917DC2D4027" pageId="56" pageNumber="1395" type="etymology">
<paragraph id="8466461AD737FF9AFF7BA917DC2D4027" blockId="56.[178,1086,219,675]" pageId="56" pageNumber="1395">E t y m o l o g y: The species epithet is the name of the island where this new species was found.</paragraph>
</subSubSection>
<subSubSection id="CCC31591D737FF9AFF7BA954DC2F40C7" pageId="56" pageNumber="1395" type="diagnosis">
<paragraph id="8466461AD737FF9AFF7BA954DC2F40C7" blockId="56.[178,1086,219,675]" pageId="56" pageNumber="1395">
C o m p a r a t i v e n o t e s: This species resembles 
<emphasis id="B6AD9A08D737FF9AFCEFA954DEB5400C" box="[806,954,595,616]" italics="true" pageId="56" pageNumber="1395">S. biliranense</emphasis>
in external characters though its body colour is notably lighter. It differs drastically by the aedeagal characters.
</paragraph>
</subSubSection>
</treatment>
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