The Genus Atheta (Coleoptera, Staphylinidae, Aleocharinae) In The Ukrainian Carpathians Author Glotov, S. State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine Author Hushtan, K. State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine & Ecological College of Lviv National Agrarian University, Zamarstynivska st., 167, Lviv, 79068 Ukraine Author Hushtan, H. State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine Author Koval, N. Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine & Uzhansky National Nature Park, Nezalezhnosti st., 7, village Velykyi Bereznyi, Ukraine Author Diedus, V. Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine text Zoodiversity 2022 56 2 91 110 http://dx.doi.org/10.15407/zoo2022.02.091 journal article 10.15407/zoo2022.02.091 2707-7268 7175709 Genus Atheta Thomson, 1858 D i a g n o s i s. Body ovoid or subparallel, rather loosely articulated, length 2.5–4.8 mm in most species; color: dark brown to black, integument usually with meshed microsculpture; pronotum with pubescence on disc forming distinct pattern, usually directed lateroposteriad from midline, but with specific patterns in different subgenera; hypomera completely visible in lateral view; mesocoxae narrowly separated; process of mesoventrite slender; tarsal formula 4–5–5; athetine bridge located in dorso-basal part of median lobe of aedeagus; spermatheca with club-shaped capsule and long stem that is narrowly looped posteriorly in most species (there are many spermathecal forms presented in Atheta ). This genus is treated here as Atheta sensu lato , and embraces a wide range of species with diverse morphology forming the Atheta complex. It includes several more or less clearly defined subgenera that are not sufficiently delimited to justify elevating them to generic status, and such an act would bring only more confusion. The subgenera of Atheta are listed below in alphabetical order because the relationships between them are not clear. For species identification, dissection and examination of genital structures is necessary (Klimaszeski et al., 2018). Subgenus Anopleta Mulsant & Rey, 1874 Atheta corvina (Thomson, 1856) Material. 1 ex. IFR.: Pistyn [Pistyń], date not specified, 1 ex. ( SMNH ) . D i s t r i b u t i o n. Europe, Siberia ( Schülke & Smetana, 2015 ); Ukrainian Carpathians: Ivano-Frankivsk Region, between Pistyn (Glotov & Hushtan, 2020). “ Poland ”, without indication of region ( Łomnicki, 1913 ) Bionomics. The species occurs in mixed and deciduous forests, in the mountains, where they live in forest litter, decaying organic residues, and in moss (Nikitskij et al., 1996; Semenov, 2007 ). Adults occur in V–VII ( Ganglbauer, 1895 ).