<document id="A8EAC1919CAFDFA5B8079EFCEA49E96B" ID-DOI="10.11646/phytotaxa.694.1.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="16723723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1754112439347" checkinUser="felipe" docAuthor="Dang, Van-Son, Le, Tuan Anh, Hoang, Thanh Son, Le, Van Son, Ha, Van Bac, Pham, Quoc Trong, Phetlasy, Souladeth, Tagane, Shuichiro &amp; Naiki, Akiyo" docDate="2025" docId="03DB87E7FFFEFFF5CBACFAA3FE5CF85B" docLanguage="en" docName="phytotaxa.694.1.057-067.pdf" docOrigin="Phytotaxa 694 (1)" docSource="https://doi.org/10.11646/phytotaxa.694.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lasianthus laoticus Tagane, Soulad. &amp; Nob. Tanaka, A. Habit, B. Branch, C. Shoot 2023" docType="treatment" docVersion="2" lastPageNumber="64" masterDocId="FFE2FF9FFFF9FFF2CB24FFD6FFF3FF94" masterDocTitle="Two new species and one new addition of Lasianthus (Rubiaceae) for the flora of Vietnam" masterLastPageNumber="67" masterPageNumber="57" pageNumber="64" updateTime="1754126368260" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="79D41A2554CCE98AE1B640BADBAE88EA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CCED1113A2515F5F2AC4AA67438522F1">
<mods:title id="22427AE42EBF41C71128C391AB3EF7E1">Two new species and one new addition of Lasianthus (Rubiaceae) for the flora of Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="4A7748771B32E9C6CC91F254F99BB861" type="personal">
<mods:role id="701910E69B8E7DBAC5FDD12226B5DF43">
<mods:roleTerm id="73CB0CC945C4663617147FB09E71326B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2268636C5594D5EA3301F1DA192E2DA">Dang, Van-Son</mods:namePart>
<mods:affiliation id="925B0431C5924862E84E4086FBFFED43">Institute of Life Sciences, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="CCEDE3770EDC48257A52ECABF4F16DAF" type="personal">
<mods:role id="320DA1C8065FD33A3752725E740E9216">
<mods:roleTerm id="9772DC1BF8DB58C5A7E358C8F4BAB90C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FDBCE36CDC2420E3AA83A9D8071412CC">Le, Tuan Anh</mods:namePart>
<mods:affiliation id="ADB6D70A7CF7FCA27DB44856CC70CC4B">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam &amp; Mien Trung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321, Huynh Thuc Khang Street, ThuaThien Hue, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="0601668860F25CC578552C74133C6526" type="personal">
<mods:role id="5209B8C8FEE163BE444A30DD74A9527D">
<mods:roleTerm id="8B4278C751BADCAE3B92918E1C8A7A2B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3616B40534F1BE4C9C455D07B22A62DD">Hoang, Thanh Son</mods:namePart>
<mods:nameIdentifier id="A3FF09C96AF80608A427DF962F2B2EEF" type="ORCID">0000-0003-0351-929X</mods:nameIdentifier>
<mods:affiliation id="C4B5A5A1A92544CB48CC7FCE4FA09925">Silviculture Research Institute, Vietnamese Academy of Forest Sciences, Ha Noi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="586AA2BE86432C212D6E9A70ACE433A2" type="email">hoangsonfsiv@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C582C12EB385C7BCABFCAB5583051AA7" type="personal">
<mods:role id="0C3DC90079BDD48B42FF781396A32603">
<mods:roleTerm id="533992DA0EE3D3E97E5C0083AFA86548">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FD77097FA6D0E8B7D0BB054A33C5C1B">Le, Van Son</mods:namePart>
<mods:nameIdentifier id="5DD19BD66904B8408A55AC4BF5D40544" type="ORCID">0000-0003-0151-0646</mods:nameIdentifier>
<mods:affiliation id="719E63BB027ACAA2DD2D1CECB4411C4D">Binh Chau-Phuoc Buu Nature Reserve, Ba Ria-Vung Tau Province, Vietnam</mods:affiliation>
<mods:nameIdentifier id="31640E2B8F33FA4774AD07A3411F8ECF" type="email">lvsonplant@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="68A2C4467C495F2434D29EAA7A053FC4" type="personal">
<mods:role id="BACF6D04182B061ADB06EF3CC65B7882">
<mods:roleTerm id="F1880C13337FA6E91E107CE157BC7C27">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83221671AC7327065164A7DE2230B3A2">Ha, Van Bac</mods:namePart>
<mods:nameIdentifier id="F6B1315E4CEA670B3012B405D8C8D1AA" type="ORCID">0009-0009-4010-7981</mods:nameIdentifier>
<mods:affiliation id="9ACDFB9A4E1115B507B1E48689501F3A">Forest Protection Department of Dakrong Nature Reserve, Quang Tri Province, Vietnam</mods:affiliation>
<mods:nameIdentifier id="9CB3D40FC64DE8BE5803687B4823DA75" type="email">vanbacha@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="ED900C6040A8A8ED204BC333BB5C3305" type="personal">
<mods:role id="8C0498F8BF0BF43B7F74F2A33AEEF853">
<mods:roleTerm id="B8351B8372FE75E40092D561A95F7F2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50FC18108ABE99BBE95639121F269DC4">Pham, Quoc Trong</mods:namePart>
<mods:nameIdentifier id="190A40278ADB6D26247F894200F9DDED" type="ORCID">0009-0004-6879-7535</mods:nameIdentifier>
<mods:affiliation id="EB0A0299CEAC758AEE694DFC1F7C2F55">Institute of Life Sciences, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="F41DF9DEF455C1163E3688A30D2873E2" type="email">phamquoctrong117@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E439663E2B235C2F023A84DCED523A5B" type="personal">
<mods:role id="A4429C1D957EA2F7AAA7E28E9DB4DC5E">
<mods:roleTerm id="89F1322431B89BA1299F754741D8FDF3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31D6C6250773A395842A1491E6441E6E">Phetlasy, Souladeth</mods:namePart>
<mods:nameIdentifier id="200562EEA96A3E9A4BB79FF627548360" type="ORCID">0000-0001-8563-1773</mods:nameIdentifier>
<mods:affiliation id="084DEC6975D1C47CDC2448CB861247C3">Faculty of Forest Science, National University of Laos, Dongdok Campus, Xaythany District, Vientiane Capital, Laos</mods:affiliation>
<mods:nameIdentifier id="26FA7ABC61A6FEF50672AB065B88C2B7" type="email">p.souladeth@nuol.edu.la</mods:nameIdentifier>
</mods:name>
<mods:name id="637F3858859086C1E4AD5DA22947CA65" type="personal">
<mods:role id="0E9A2A10153AD426EF9B6A6A5C3D90E6">
<mods:roleTerm id="EDBA43AEE43F846313408C2B74F7FA48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EDC94EB51E1A4AC26B602991EEF65408">Tagane, Shuichiro</mods:namePart>
<mods:nameIdentifier id="479A742760A461428564C24F2EC347A5" type="ORCID">0000-0002-1974-7329</mods:nameIdentifier>
<mods:affiliation id="189DAD93FB1B121F816CEB3F34C3BB11">The Kagoshima University Museum, Kagoshima University, 1 - 21 - 30, Korimoto, Kagoshima, 890 - 0065, Japan</mods:affiliation>
<mods:nameIdentifier id="72A1FCB5ADC320A23066059FD1211336" type="email">stagane29@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="ADBA0C78E8876D045976949E9B35F5CC" type="personal">
<mods:role id="5799C91E31452C4BA50B73D9FD443937">
<mods:roleTerm id="73F290D50C977270485E387E3B822A2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1B4D9157E489A35704FB62A7BAEB04A">Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier id="A12D8871F8D34B7396C9F7CC78BD23F8" type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation id="6773CFCA5D8D7DD06225FC16BC5BD924">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan</mods:affiliation>
<mods:nameIdentifier id="638F010C8157DB1D6A422C3EEE407BC1" type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5A53BCE0840E51D64337D326079F5C13">text</mods:typeOfResource>
<mods:relatedItem id="45E2D3F82AF87D7C79E1D22EFF31B233" type="host">
<mods:titleInfo id="763763E958F8C7BEC2B17039D099B155">
<mods:title id="C84C0AABB14CCAB9777566AF982DFCB6">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="E99F87486C0889B7A50C3F91DA57B147">
<mods:date id="C63FC13B65A28B3A5630AED3F41EC069">2025</mods:date>
<mods:detail id="2F9BFB907D4ABC3C7CDBD09A2093C965" type="pubDate">
<mods:number id="82B6EBD467CC49E6D7DA1D237527F867">2025-03-18</mods:number>
</mods:detail>
<mods:detail id="9489884F37414C3C76A27254B51FB320" type="volume">
<mods:number id="73ACA425F9DA31710983CA50E2C1046A">694</mods:number>
</mods:detail>
<mods:detail id="9E90DB56FC6F56DA00AD158502E7C984" type="issue">
<mods:number id="25E2E3BA6FB521C998C336159DF99CBE">1</mods:number>
</mods:detail>
<mods:extent id="D2E6136D693F37B757E7498F27A65AB7" unit="page">
<mods:start id="D1E0D04E7B9F2A882A71454D9CC8E883">57</mods:start>
<mods:end id="90D4A7806B5C0AD742C169025537924A">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="7B3EFF11D892D7891BEA323B3DA0F34E">
<mods:url id="FB81AF3C0F61AF1E83114AE26B87D95C">https://doi.org/10.11646/phytotaxa.694.1.4</mods:url>
</mods:location>
<mods:classification id="B33253E1938CFC8357D1640C440FFF14">journal article</mods:classification>
<mods:identifier id="2B82D22765F39653C7F70D9D3C34C995" type="DOI">10.11646/phytotaxa.694.1.4</mods:identifier>
<mods:identifier id="5761617574B8ECC5275E62878F8D6F6E" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="212C62212142EFC828D1D0B828C70FF0" type="Zenodo-Dep">16723723</mods:identifier>
</mods:mods>
<treatment id="03DB87E7FFFEFFF5CBACFAA3FE5CF85B" ID-DOI="http://doi.org/10.5281/zenodo.16723717" ID-Zenodo-Dep="16723717" LSID="urn:lsid:plazi:treatment:03DB87E7FFFEFFF5CBACFAA3FE5CF85B" httpUri="http://treatment.plazi.org/id/03DB87E7FFFEFFF5CBACFAA3FE5CF85B" lastPageNumber="64" pageId="7" pageNumber="64">
<subSubSection id="C368657AFFFEFFF5CBACFAA3FBF9FA1B" box="[136,1034,1397,1424]" pageId="7" pageNumber="64" type="nomenclature">
<paragraph id="8BCD36F1FFFEFFF5CBACFAA3FBF9FA1B" blockId="7.[136,1034,1397,1424]" box="[136,1034,1397,1424]" pageId="7" pageNumber="64">
<heading id="D085819DFFFEFFF5CBACFAA3FBF9FA1B" box="[136,1034,1397,1424]" fontSize="11" level="2" pageId="7" pageNumber="64" reason="3">
<taxonomicName id="4C724D72FFFEFFF5CBACFAA3FCADFA1B" ID-CoL="CDJ5P" authority="Tagane, Soulad. &amp; Nob. Tanaka (2023: 148)" authorityName="Tagane, Soulad. &amp; Nob. Tanaka, A. Habit, B. Branch, C. Shoot" authorityPageNumber="148" authorityYear="2023" box="[136,862,1397,1424]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="64" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis id="B906EAE3FFFEFFF5CBACFAA3FE99FA04" bold="true" box="[136,362,1397,1424]" italics="true" pageId="7" pageNumber="64">Lasianthus laoticus</emphasis>
Tagane, Soulad. &amp; Nob.Tanaka (2023: 148)
</taxonomicName>
. 
<figureCitation id="13492A74FFFEFFF5C84DFAA3FC21FA04" box="[873,978,1397,1424]" captionStart="FIGURE 5" captionStartId="8.[136,229,2022,2044]" captionTargetBox="[184,1403,187,1984]" captionTargetId="figure-21@8.[184,1403,187,1995]" captionTargetPageId="8" captionText="FIGURE 5. Specimen of Lasianthus laoticus Tagane, Soulad. &amp; Nob.Tanaka (Dang 664, VNM). Photoed by Van-Son Dang." figureDoi="http://doi.org/10.5281/zenodo.16723735" httpUri="https://zenodo.org/record/16723735/files/figure.png" pageId="7" pageNumber="64">Figures 5</figureCitation>
&amp; 
<figureCitation id="13492A74FFFEFFF5C8D1FAA3FBF5FA1B" box="[1013,1030,1397,1423]" captionStart="FIGURE 6" captionStartId="9.[136,229,1987,2009]" captionTargetBox="[194,1394,187,1960]" captionTargetId="figure-19@9.[194,1394,187,1960]" captionTargetPageId="9" captionText="FIGURE 6. Lasianthus laoticus Tagane, Soulad. &amp; Nob.Tanaka, A. Habit, B. Branch, C. Shoot apex showing expanding young leaves, D. Stipule, E. Abaxial leaf surface, F–G. Inflorescences and flowers, H. Young fruit, I. Mature fruit. Photoed by Van-Son Dang." figureDoi="http://doi.org/10.5281/zenodo.16723737" httpUri="https://zenodo.org/record/16723737/files/figure.png" pageId="7" pageNumber="64">6</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368657AFFFEFFF5CBACFA17FDDBF98A" pageId="7" pageNumber="64" type="materials_examined">
<paragraph id="8BCD36F1FFFEFFF5CBACFA17FDDBF98A" blockId="7.[136,1452,1472,1567]" pageId="7" pageNumber="64">
<collectionCode id="ED63AE34FFFEFFF5CBACFA17FF34FA42" box="[136,199,1473,1494]" pageId="7" pageNumber="64">TYPE</collectionCode>
:— 
<collectingCountry id="F3657661FFFEFFF5CBC0FA17FED6FA42" box="[228,293,1473,1494]" name="Laos" pageId="7" pageNumber="64">LAOS</collectingCountry>
. 
<collectingRegion id="49B6F813FFFEFFF5CA08FA17FE24FA43" box="[300,471,1473,1495]" country="Laos" name="Attapu" pageId="7" pageNumber="64">Attapeu Province</collectingRegion>
: Sanamxai District, roadside, at the edge of evergreen forests, 
<geoCoordinate id="EE465036FFFEFFF5CF09FA16FB33FA43" box="[1069,1216,1472,1495]" degrees="14" direction="north" minutes="47" orientation="latitude" pageId="7" pageNumber="64" precision="1" seconds="18.18" value="14.7883835">14°47′18.18″N</geoCoordinate>
, 
<geoCoordinate id="EE465036FFFEFFF5CFEDFA16FA96FA43" box="[1225,1381,1472,1495]" degrees="106" direction="east" minutes="25" orientation="longitude" pageId="7" pageNumber="64" precision="1" seconds="24.90" value="106.423584">106°25′24.90″E</geoCoordinate>
, 
<quantity id="4C8A9B14FFFEFFF5CE4BFA17FA5FFA42" box="[1391,1452,1473,1494]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="7" pageNumber="64" unit="m" value="183.0">183 m</quantity>
elevation, 
<date id="FFCC1031FFFEFFF5CA05FA33FE62FA6E" box="[289,401,1509,1531]" pageId="7" pageNumber="64" value="2019-07-06">6 July 2019</date>
, 
<emphasis id="B906EAE3FFFEFFF5CABEFA30FAF8FA6E" box="[410,1291,1509,1531]" italics="true" pageId="7" pageNumber="64">
<collectionCode id="ED63AE34FFFEFFF5CABEFA30FE5AFA6F" box="[410,425,1510,1531]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="7" pageNumber="64" type="Herbarium">P</collectionCode>
. Souladeth, 
<collectionCode id="ED63AE34FFFEFFF5C938FA33FDD8FA6E" box="[540,555,1509,1530]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="7" pageNumber="64" type="Herbarium">S</collectionCode>
. Tagane, 
<collectionCode id="ED63AE34FFFEFFF5C9A0FA30FD64FA6F" box="[644,663,1510,1531]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="7" pageNumber="64" type="Herbarium">N</collectionCode>
. Tanaka, 
<collectionCode id="ED63AE34FFFEFFF5C9D4FA30FCF0FA6F" box="[752,771,1510,1531]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="7" pageNumber="64" type="Herbarium">K</collectionCode>
. Souttakone, 
<collectionCode id="ED63AE34FFFEFFF5C8A5FA30FC7CFA6F" box="[897,911,1510,1531]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="7" pageNumber="64" type="Herbarium">Y</collectionCode>
. Suyama, 
<collectionCode id="ED63AE34FFFEFFF5C8CBFA30FBF1FA6F" box="[1007,1026,1510,1531]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="7" pageNumber="64" type="Herbarium">K</collectionCode>
. Phengmala, 
<collectionCode id="ED63AE34FFFEFFF5CFA7FA30FB65FA6F" box="[1155,1174,1510,1531]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="7" pageNumber="64" type="Herbarium">N</collectionCode>
. Ishii 
<collectionCode id="ED63AE34FFFEFFF5CFEAFA33FB2FFA6E" box="[1230,1244,1509,1530]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="7" pageNumber="64" type="Herbarium">L</collectionCode>
3024
</emphasis>
(
<typeStatus id="54C98853FFFEFFF5CE33FA33FA9EFA6F" box="[1303,1389,1509,1531]" pageId="7" pageNumber="64" type="holotype">holotype</typeStatus>
<collectionCode id="ED63AE34FFFEFFF5CE55FA33FA57FA6E" box="[1393,1444,1509,1530]" pageId="7" pageNumber="64">FOF</collectionCode>
!, 
<typeStatus id="54C98853FFFEFFF5CB99F9DFFEFFF98B" box="[189,268,1545,1567]" pageId="7" pageNumber="64" type="isotype">isotypes</typeStatus>
<collectionCode id="ED63AE34FFFEFFF5CA36F9DFFEB5F98A" box="[274,326,1545,1566]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="7" pageNumber="64" type="Herbarium">KAG</collectionCode>
[
<collectionCode id="ED63AE34FFFEFFF5CA76F9DFFE79F98A" box="[338,394,1545,1566]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="7" pageNumber="64" type="Herbarium">KAG</collectionCode>
129297], 
<collectionCode id="ED63AE34FFFEFFF5CAC0F9DFFDEAF98A" box="[484,537,1545,1566]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="7" pageNumber="64" type="Herbarium">TNS</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="C368657AFFFEFFF5CBACF99BFDC7F8F7" pageId="7" pageNumber="64" type="distribution">
<paragraph id="8BCD36F1FFFEFFF5CBACF99BFD0DF924" blockId="7.[136,1452,1613,1999]" pageId="7" pageNumber="64">
<emphasis id="B906EAE3FFFEFFF5CBACF99BFDB2F9F3" bold="true" box="[136,577,1613,1639]" pageId="7" pageNumber="64">Distribution, habitat and phenology:</emphasis>
— 
<collectingCountry id="F3657661FFFEFFF5C97AF998FD67F9FC" box="[606,660,1614,1640]" name="Laos" pageId="7" pageNumber="64">Laos</collectingCountry>
(
<collectingRegion id="49B6F813FFFEFFF5C987F99BFD0DF9F3" box="[675,766,1613,1639]" country="Laos" name="Attapu" pageId="7" pageNumber="64">Attapeu</collectingRegion>
and 
<collectingRegion id="49B6F813FFFEFFF5C813F99BFC49F9F3" box="[823,954,1613,1639]" country="Laos" name="Champasak" pageId="7" pageNumber="64">Champasak</collectingRegion>
Provinces) and 
<collectingCountry id="F3657661FFFEFFF5CF56F99BFB22F9F3" box="[1138,1233,1613,1639]" name="Vietnam" pageId="7" pageNumber="64">Vietnam</collectingCountry>
(Binh Chau-Phuoc Buu Nature Reserve, 
<collectingRegion id="49B6F813FFFEFFF5CA5AF9A4FD46F91F" box="[382,693,1649,1676]" country="Vietnam" name="Ba Ria-Vung Tau" pageId="7" pageNumber="64">Ba Ria-Vung Tau Province</collectingRegion>
). It grows in lowland evergreen forests, 
<quantity id="4C8A9B14FFFEFFF5CFA5F9A7FB07F918" box="[1153,1268,1649,1676]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.9" metricValueMin="0.2" pageId="7" pageNumber="64" unit="m" value="105.0" valueMax="190.0" valueMin="20.0">20–190 m</quantity>
elev. Flowering and fruiting specimens were collected in July to August.
</paragraph>
<paragraph id="8BCD36F1FFFEFFF5CB99F96FFD1CF947" blockId="7.[136,1452,1613,1999]" box="[189,751,1721,1750]" pageId="7" pageNumber="64">
<emphasis id="B906EAE3FFFEFFF5CB99F96FFE66F947" bold="true" box="[189,405,1721,1747]" pageId="7" pageNumber="64">Vernacular name:</emphasis>
—Xú hƯƠng lào (Vietnamese).
</paragraph>
<paragraph id="8BCD36F1FFFEFFF5CB99F90BFDC7F8F7" blockId="7.[136,1452,1613,1999]" pageId="7" pageNumber="64">
<emphasis id="B906EAE3FFFEFFF5CB99F90BFEFAF963" bold="true" box="[189,265,1757,1783]" pageId="7" pageNumber="64">Notes:</emphasis>
— 
<taxonomicName id="4C724D72FFFEFFF5CA02F90BFE0CF963" authorityName="Tagane, Soulad. &amp; Nob. Tanaka, A. Habit, B. Branch, C. Shoot" authorityYear="2023" box="[294,511,1757,1783]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="64" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis id="B906EAE3FFFEFFF5CA02F90BFE0CF963" box="[294,511,1757,1783]" italics="true" pageId="7" pageNumber="64">Lasianthus laoticus</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3657661FFFEFFF5C9C5F908FCE4F96C" box="[737,791,1758,1784]" name="Laos" pageId="7" pageNumber="64">Laos</collectingCountry>
by 
<bibRefCitation id="EFE34B00FFFEFFF5C81AF908FBE9F963" author="Tagane, S. &amp; Souladeth, P. &amp; Tanaka, N. &amp; Phengmala, K. &amp; Yahara, T." box="[830,1050,1757,1784]" pageId="7" pageNumber="64" pagination="145 - 153" refId="ref4949" refString="Tagane, S., Souladeth, P., Tanaka, N., Phengmala, K. &amp; Yahara, T. (2023) Two new species and a new record of Lasianthus (Rubiaceae) in Laos. Phytotaxa 585 (2): 145-153. https://doi.org/10.11646/phytotaxa.585.2.5" type="journal article" year="2023">
Tagane 
<emphasis id="B906EAE3FFFEFFF5C8B0F908FC38F963" box="[916,971,1757,1783]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
(2023)
</bibRefCitation>
based on two collections, 
<emphasis id="B906EAE3FFFEFFF5CE1FF90BFEFDF88F" italics="true" pageId="7" pageNumber="64">Souladeth et al. L3024</emphasis>
and 
<emphasis id="B906EAE3FFFEFFF5CA66F8D7FE79F88F" box="[322,394,1793,1819]" italics="true" pageId="7" pageNumber="64">L2222</emphasis>
. This species is easily recognizable in the field in having brownish hairy branches, cordate leaf base, secondary veins 8–11 pairs, long peduncles 
<quantity id="4C8A9B14FFFEFFF5C991F8F3FCEEF8D4" box="[693,797,1829,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="7" pageNumber="64" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
long, flowers white with a faint tinge of yellow, and fruits subglobose or ovoid, black when ripe.
</paragraph>
</subSubSection>
<subSubSection id="C368657AFFFEFFF5CB99F8BBFE5CF85B" pageId="7" pageNumber="64" type="description">
<paragraph id="8BCD36F1FFFEFFF5CB99F8BBFE5CF85B" blockId="7.[136,1452,1613,1999]" pageId="7" pageNumber="64">
<emphasis id="B906EAE3FFFEFFF5CB99F8BBFE4AF813" bold="true" box="[189,441,1901,1927]" pageId="7" pageNumber="64">Specimen examined:</emphasis>
— 
<collectingCountry id="F3657661FFFEFFF5CAF2F8B8FDA9F81C" box="[470,602,1902,1928]" name="Vietnam" pageId="7" pageNumber="64">VIETNAM</collectingCountry>
. 
<collectingRegion id="49B6F813FFFEFFF5C94EF8B8FC58F813" box="[618,939,1901,1928]" country="Vietnam" name="Ba Ria-Vung Tau" pageId="7" pageNumber="64">Ba Ria-Vung Tau Province</collectingRegion>
: Binh Chau-Phuoc Buu Nature Reserve, in evergreen forests, 
<geoCoordinate id="EE465036FFFEFFF5CA78F845FDF4F83A" box="[348,519,1939,1966]" degrees="10" direction="north" minutes="35" orientation="latitude" pageId="7" pageNumber="64" precision="1" seconds="12.04" value="10.586678">10°35′12.04″N</geoCoordinate>
, 
<geoCoordinate id="EE465036FFFEFFF5C931F845FD38F83A" box="[533,715,1939,1966]" degrees="107" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="64" precision="1" seconds="25.00" value="107.556946">107°33′25.00″E</geoCoordinate>
, 
<quantity id="4C8A9B14FFFEFFF5C9FDF847FCE6F838" box="[729,789,1937,1964]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="64" unit="m" value="28.0">28 m</quantity>
elev., 
<date id="FFCC1031FFFEFFF5C847F847FBEDF83F" box="[867,1054,1937,1964]" pageId="7" pageNumber="64" value="2022-08-16">16 August 2022</date>
, 
<emphasis id="B906EAE3FFFEFFF5CF08F844FA5FF83F" box="[1068,1452,1937,1964]" italics="true" pageId="7" pageNumber="64">V.-S. Dang &amp; L.V. Son Dang-664</emphasis>
(VNM [VNM00070336]).
</paragraph>
</subSubSection>
</treatment>
</document>