<document id="D5E33EDC979756171991385669B582E7" ID-CLB-Dataset="41300" ID-DOI="10.5281/zenodo.186351" ID-GBIF-Dataset="bb17e039-5346-4e01-a9ba-59b87e9e6a4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="186351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050352768" checkinUser="plazi" docAuthor="Maus, Mariana Chani-Posse De" docDate="2009" docId="03D787C9FFAEFF9FFF1BCBAFCBC8E33D" docLanguage="en" docName="zt02034p042.pdf" docOrigin="Zootaxa 2034" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gabrius tucumanensis Bernhauer 1927, comb. nov." docType="treatment" docVersion="11" lastPageNumber="34" masterDocId="FFEEFFB1FFACFF9CFF8CCD70CA68E133" masterDocTitle="Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes" masterLastPageNumber="42" masterPageNumber="31" pageNumber="33" updateTime="1732667960883" updateUser="ExternalLinkService">
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<mods:title id="3C4E3040C717CC4903978DA1304F89C7">Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes</mods:title>
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<mods:namePart id="E56CF492E853B938ECD08D0B3B1400E3">Maus, Mariana Chani-Posse De</mods:namePart>
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<subSubSection id="C3646554FFAEFF9EFF1BCBAFC94DE7CA" box="[151,805,1759,1785]" pageId="2" pageNumber="33" type="nomenclature">
<paragraph id="8BC136DFFFAEFF9EFF1BCBAFC94DE7CA" blockId="2.[151,805,1759,1785]" box="[151,805,1759,1785]" pageId="2" pageNumber="33">
<heading id="D08981B3FFAEFF9EFF1BCBAFC94DE7CA" bold="true" box="[151,805,1759,1785]" fontSize="11" level="1" pageId="2" pageNumber="33" reason="1">
<emphasis id="B90AEACDFFAEFF9EFF1BCBAFC94DE7CA" bold="true" box="[151,805,1759,1785]" pageId="2" pageNumber="33">
<taxonomicName id="4C7E4D5CFFAEFF9EFF1BCBAFC8F9E7CA" ID-CoL="6K78G" authority="Bernhauer, 1927" authorityName="Bernhauer" authorityYear="1927" box="[151,657,1759,1785]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="tucumanensis" status="comb. nov.">
<emphasis id="B90AEACDFFAEFF9EFF1BCBAFCBC0E7CA" bold="true" box="[151,424,1759,1785]" italics="true" pageId="2" pageNumber="33">Gabrius tucumanensis</emphasis>
(
<bibRefCitation id="EFEF4B2EFFAEFF9EFE3BCBAFC8E0E7CA" author="Bernhauer" box="[439,648,1759,1785]" pageId="2" pageNumber="41" refString="Bernhauer, M. (1927) Zur Staphylinidenfauna Sud-Amerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte, (A) 91 (12) (1925), 229 - 264." type="journal article" year="1927">Bernhauer, 1927</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="A23957B6FFAEFF9EFD12CBAFC94DE7CA" box="[670,805,1759,1785]" pageId="2" pageNumber="33">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAEFF9EFF1BCA59C93CE653" pageId="2" pageNumber="33" type="reference_group">
<paragraph id="8BC136DFFFAEFF9EFF1BCA59C93CE653" blockId="2.[151,1436,1833,1888]" pageId="2" pageNumber="33">
<treatmentCitationGroup id="AB6E11F1FFAEFF9EFF1BCA59C93CE653" pageId="2" pageNumber="33">
<taxonomicName id="4C7E4D5CFFAEFF9EFF1BCA59C8C3E673" ID-CoL="4GC2F" authority="Bernhauer, 1927: 245" authorityName="Bernhauer" authorityPageNumber="245" authorityYear="1927" box="[151,683,1833,1856]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAEFF9EFF1BCA59CBC0E673" box="[151,424,1833,1856]" italics="true" pageId="2" pageNumber="33">Philonthus tucumanensis</emphasis>
<treatmentCitation id="0ADF10CEFFAEFF9EFE39CA59C8C3E673" author="Bernhauer" box="[437,683,1833,1856]" page="245" pageId="2" pageNumber="33" year="1927">
<bibRefCitation id="EFEF4B2EFFAEFF9EFE39CA59C807E673" author="Bernhauer" box="[437,623,1833,1856]" pageId="2" pageNumber="41" refString="Bernhauer, M. (1927) Zur Staphylinidenfauna Sud-Amerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte, (A) 91 (12) (1925), 229 - 264." type="journal article" year="1927">Bernhauer, 1927</bibRefCitation>
: 245
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0ADF10CEFFAEFF9EFD30CA59C9A2E673" author="Scheerpeltz" box="[700,970,1833,1856]" page="1365" pageId="2" pageNumber="33" year="1933">
<bibRefCitation id="EFEF4B2EFFAEFF9EFD30CA59C9EDE673" author="Scheerpeltz" box="[700,901,1833,1856]" pageId="2" pageNumber="42" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1365
</treatmentCitation>
(catalog); 
<treatmentCitation id="0ADF10CEFFAEFF9EFBC7CA59CF3EE673" author="Blackwelder" box="[1099,1366,1833,1856]" page="135" pageId="2" pageNumber="33" year="1944">
<bibRefCitation id="EFEF4B2EFFAEFF9EFBC7CA59CF75E673" author="Blackwelder" box="[1099,1309,1833,1856]" pageId="2" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 135
</treatmentCitation>
(list); 
<treatmentCitation id="0ADF10CEFFAEFF9EFF4BCA39CBCAE653" author="Herman" box="[199,418,1865,1888]" httpUri="http://treatment.plazi.org/id/03C1F679FDE9FDEEFF2FFF7B825AF912" page="2982" pageId="2" pageNumber="33" year="2001">
<bibRefCitation id="EFEF4B2EFFAEFF9EFF4BCA39CB0BE653" author="Herman" box="[199,355,1865,1888]" pageId="2" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2982
</treatmentCitation>
(catalog); Chani-Posse, 2004: 231 (list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAEFF9FFF1BCAFFCE4DE0F3" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33" type="materials_examined">
<paragraph id="8BC136DFFFAEFF9FFF1BCAFFCF4EE181" blockId="2.[151,1437,1935,2000]" lastBlockId="3.[151,1437,152,526]" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33">
<emphasis id="B90AEACDFFAEFF9EFF1BCAFFCB84E69A" bold="true" box="[151,492,1935,1961]" pageId="2" pageNumber="33">
<typeStatus id="54C5887DFFAEFF9EFF1BCAFFCAB1E69A" box="[151,217,1935,1961]" pageId="2" pageNumber="33">Type</typeStatus>
material. 
<typeStatus id="54C5887DFFAEFF9EFEEACAFFCB84E69A" box="[358,492,1935,1961]" pageId="2" pageNumber="33" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated; 
<figureCitation id="13452A5AFFAEFF9EFD56CAE0C95AE699" box="[730,818,1936,1962]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="2" pageNumber="33">Figs. 3</figureCitation>
a–b), Ƥ, “ 
<collectingCountry id="F369764FFFAEFF9EFC35CAE0CE57E699" box="[953,1087,1936,1962]" name="Saint Pierre and Miquelon" pageId="2" pageNumber="33">San Pedro</collectingCountry>
Colalao / Tucum. Weiser”, “ 
<taxonomicName id="4C7E4D5CFFAEFF9EFF28CAC7CBC9E6E3" authority="Bernh." authorityName="Bernh." box="[164,417,1974,2000]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAEFF9EFF28CAC7CB2FE6E3" box="[164,327,1975,2000]" italics="true" pageId="2" pageNumber="33">tucumanensis</emphasis>
Bernh.
</taxonomicName>
/ 
<typeStatus id="54C5887DFFAEFF9EFE36CAC6C86BE6E3" box="[442,515,1974,2000]" pageId="2" pageNumber="33">Typus</typeStatus>
don. Bruch” (yellow label), “Chicago NHMus. / M. Bernhauer Collection”, “ 
<typeStatus id="54C5887DFFAFFF9FFF2DCDE8CB71E181" box="[161,281,152,178]" pageId="3" pageNumber="34" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFAFFF9FFEACCDE8C978E181" authority="Bernhauer, 1927" authorityName="Bernhauer" authorityYear="1927" box="[288,784,152,178]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAFFF9FFEACCDE8C82EE182" box="[288,582,152,177]" italics="true" pageId="3" pageNumber="34">Philonthus tucumanensis</emphasis>
<bibRefCitation id="EFEF4B2EFFAFFF9FFDC1CDE8C978E181" author="Bernhauer" box="[589,784,152,178]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1927) Zur Staphylinidenfauna Sud-Amerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte, (A) 91 (12) (1925), 229 - 264." type="journal article" year="1927">Bernhauer, 1927</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFAFFF9FFB33CDE8CF73E181" LSID="urn:lsid:biocol.org:col:34795" box="[1215,1307,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="34">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFAFFF9FFF4ACDCEC925E015" blockId="3.[151,1437,152,526]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF4ACDCEC977E1EB" bold="true" box="[198,799,190,216]" pageId="3" pageNumber="34">
Additional material. 
<collectingCountry id="F369764FFFAFFF9FFE5BCDCEC8E7E1EB" box="[471,655,190,216]" name="Argentina" pageId="3" pageNumber="34">ARGENTINA</collectingCountry>
: San Luis:
</emphasis>
2533, 4ƤƤ, Irideus, Quebrada del Tala, 
<quantity id="4C869B3AFFAFFF9FFABBCDCECFFFE1EB" box="[1335,1431,190,216]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="34" unit="m" value="1100.0">1100 m</quantity>
, 
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, 
<geoCoordinate id="EE4A5018FFAFFF9FFEA5CD95CBD1E1CC" box="[297,441,229,255]" direction="west" orientation="longitude" pageId="3" pageNumber="34" precision="15" value="-64.95638">64°57'23''W</geoCoordinate>
, litter in moist narrow quebrada, 
<date id="FFC0101FFFAFFF9FFCB3CD95C9AAE1CC" box="[831,962,229,255]" pageId="3" pageNumber="34" value="2000-12-08">8.XII.2000</date>
, L. Herman (
<collectionCode id="ED6FAE1AFFAFFF9FFBD2CD95CED6E1CC" LSID="urn:lsid:biocol.org:col:34925" box="[1118,1214,229,255]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="34">AMNH</collectionCode>
); 
<emphasis id="B90AEACDFFAFFF9FFB5FCD95CF27E1CC" bold="true" box="[1235,1359,229,255]" pageId="3" pageNumber="34">Tucumán:</emphasis>
3, Ƥ, Villa Nougués, 
<quantity id="4C869B3AFFAFFF9FFEC2CC7CCBC1E015" box="[334,425,268,294]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="34" unit="m" value="1350.0">1350 m</quantity>
, 
<date id="FFC0101FFFAFFF9FFE39CC7CC82DE015" box="[437,581,268,294]" pageId="3" pageNumber="34" value="1971-12-11">11.XII.1971</date>
, L. Herman (
<collectionCode id="ED6FAE1AFFAFFF9FFD6CCC7CC929E015" LSID="urn:lsid:biocol.org:col:34925" box="[736,833,268,294]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="34">AMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFAFFF9FFF4ACC42CE4DE0F3" blockId="3.[151,1437,152,526]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF4ACC42CB7CE07F" bold="true" box="[198,276,306,332]" pageId="3" pageNumber="34">Notes.</emphasis>
I designate the 
<typeStatus id="54C5887DFFAFFF9FFE5ECC42C855E07F" box="[466,573,306,332]" pageId="3" pageNumber="34" type="lectotype">lectotype</typeStatus>
rather than assume that the above specimen is the 
<typeStatus id="54C5887DFFAFFF9FFB11CC42CF6FE07F" box="[1181,1287,306,332]" pageId="3" pageNumber="34" type="holotype">holotype</typeStatus>
, since in the original description 
<bibRefCitation id="EFEF4B2EFFAFFF9FFE1BCC29C81DE040" author="Bernhauer" box="[407,629,345,371]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1927) Zur Staphylinidenfauna Sud-Amerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte, (A) 91 (12) (1925), 229 - 264." type="journal article" year="1927">Bernhauer (1927)</bibRefCitation>
gave no information about the number of specimens examined. Therefore, it is possible that this name was based on more than one specimen (
<bibRefCitation id="EFEF4B2EFFAFFF9FFBB9CCF0CAA0E0F3" author="ICZN" pageId="3" pageNumber="42" refString="ICZN (1999) International Code of Zoological Nomenclatur e. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">
<collectionCode id="ED6FAE1AFFAFFF9FFBB9CCF0CE13E0A9" box="[1077,1147,384,410]" pageId="3" pageNumber="34">ICZN</collectionCode>
1999, Recommendation 73F
</bibRefCitation>
). The 
<typeStatus id="54C5887DFFAFFF9FFE9ECCD6CB15E0F3" box="[274,381,422,448]" pageId="3" pageNumber="34" type="lectotype">lectotype</typeStatus>
agrees with the original description by 
<bibRefCitation id="EFEF4B2EFFAFFF9FFCC2CCD6CE49E0F3" author="Bernhauer" box="[846,1057,422,448]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1927) Zur Staphylinidenfauna Sud-Amerikas, insbesondere Argentiniens. Archiv fur Naturgeschichte, (A) 91 (12) (1925), 229 - 264." type="journal article" year="1927">Bernhauer (1927)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAFFF9FFF4ACCBDCBC8E33D" pageId="3" pageNumber="34" type="discussion">
<paragraph id="8BC136DFFFAFFF9FFF4ACCBDCBC8E33D" blockId="3.[151,1437,152,526]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF4ACCBDCB3DE0D4" bold="true" box="[198,341,461,487]" pageId="3" pageNumber="34">Comments.</emphasis>
Arguments supporting the transfer of 
<taxonomicName id="4C7E4D5CFFAFFF9FFC9ACCBDC9BBE0D5" box="[790,979,461,486]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAFFF9FFC9ACCBDC9BBE0D5" box="[790,979,461,486]" italics="true" pageId="3" pageNumber="34">P. tucumanensis</emphasis>
</taxonomicName>
to 
<taxonomicName id="4C7E4D5CFFAFFF9FFC74CCBDCE31E0D5" box="[1016,1113,461,486]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFC74CCBDCE31E0D5" box="[1016,1113,461,486]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
are given in Discussion for 
<emphasis id="B90AEACDFFAFFF9FFF1BCC84CB7BE33E" box="[151,275,500,525]" italics="true" pageId="3" pageNumber="34">G. n i d i c o l a</emphasis>
, see below.
</paragraph>
</subSubSection>
</treatment>
</document>