Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina) Author Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H. text Bulletin of the American Museum of Natural History 2001 2001-07-18 2001 265 2441 3020 http://dx.doi.org/10.1206/0003-0090.265.1.5 journal article 10.1206/0003-0090.265.1.5 0003-0090 13239102 longicollis Bernhauer, 1908 , see: Bisnius . longicornis Stephens, 1832: 237 ( Philonthus ; Type locality: London). — Curtis, 1829: 26 ( Philonthus ; nomen nudum; catalog; Britain). — Erichson, 1839a: 466 ( Philonthus ; [ Note : Bernhauer and Schubert, 1914: 345 cited varians variety b as a synonym of longicornis ; Erichson included it under opacus , but the variety is unavailable]; characters). — Stephens, 1839: 397 ( Philonthus ; characters; England ). — Fairmaire, 1849: 290 ( Staphylinus ; [ Note: Misidentification: Bernhauer and Schubert, 1914: 345 cited varians Fairmaire as a synonym of longicornis ; Fairmaire did not describe the species as new, he attributed it to Paykull]; notes; Tahiti). — Hardy, 1851: 29 ( Philonthus ; synonym of varians ). — Fauvel, 1874: 254 [= 1874a: 480] ( Philonthus ; characters; France ; Mediterranean region; Siberia ; Japan ; Ceylon ; Azores; Canary Islands; Madeira; Cape Verde Islands; Ascension Island ; South Africa ; Madagascar ; Tahiti; Australia ; North and South America). — Solsky, 1874: 178 ( Philonthus ; characters; Turkestan). — J. Sahlberg, 1876: 20 ( Philonthus ; Finland ). — Wollaston, 1877: 28 ( Philonthus ; characters; St. Helena ). — Fauvel, 1877: 264 ( Philonthus ; characters; Australia ; Tahiti). — Fauvel, 1878: 125 [= 1878a: 45] ( Philonthus ; North Africa). — Fauvel, 1878g : 548 ( Philonthus ; New Caledonia ). — Horn, 1884: 195 ( Philonthus ; characters; North America). — Sharp, 1885: 416 ( Philonthus ; Mexico ). — Fauvel, 1886: 56 [= 1886a: 48] ( Philonthus ; North Africa). — Fowler, 1888: 273 ( Philonthus ; characters; Britain). — Fauvel, 1889b: 266 ( Philonthus ; New Caledonia ). — Fauvel, 1891: 115 ( Philonthus ; Venezuela ). — Ganglbauer, 1895: 454 ( Philonthus ; characters; Europe). — Fauvel, 1902b: 112 ( Philonthus ; Barbary; Egypt ; Azores; Madeira; Canaries). — Bernhauer, 1902b: 27 ( Philonthus ; Ceylon ). — Fauvel, 1903: 159 ( Philonthus ; India ). — Fauvel, 1903d: 271 ( Philonthus ; New Caledonia ; Iles des Pins). — Fauvel, 1904a: 59 ( Philonthus ; Nilghiris). — Fall and Cockerell, 1907: 166 ( Philonthus ; New Mexico ). — Bernhauer, 1908: 19 ( Philonthus ; Australia ). — Bernhauer, 1908c: 110 ( Philonthus ; German Southwest Africa ). — Reitter, 1909: 130 ( Philonthus ; characters; Germany ). — Blatchley, 1910: 383 ( Philonthus ; characters; Indiana ). — Eichelbaum, 1913: 130 ( Philonthus ; Tanzania ). — Bernhauer and Schubert, 1914: 344 ( Philonthus ; catalog). — Johansen, 1914: 408 ( Philonthus ; characters; Denmark ). — Bernhauer, 1920: 16 ( Philonthus ; Australia ). — Gridelli, 1920a: 18, 19 ( Philonthus ; characters; Italy ). — Bernhauer, 1921a: 42 ( Philonthus ; Easter Island). — Bernhauer, 1922c: 232 ( Philonthus ; Formosa ). — Cameron, 1924: 87 ( Philonthus ; New Guinea ). — Dodero, 1925: 5 ( Philonthus ; Libya ). — Lea, 1925: 220 ( Philonthus ; Australian introduction). — Portevin, 1929: 363 ( Philonthus ; characters; France ). — Gridelli, 1930: 74 ( Philonthus ; Libya ). — Bernhauer, 1931d: 587 ( Philonthus ; Abyssinia ). — Cameron, 1932: 127 ( Philonthus ; characters; Ceylon ; India ). — Scheerpeltz, 1933: 1349 ( Philonthus ; catalog).

Bulletin AMNH STAPHYLININAE : PHILONTHUS 2867
— Cameron, 1933e: 82 ( Philonthus ; Society Islands).
— Cameron, 1933f: 80 ( Philonthus ; Marquesas Islands).
— Wüsthoff, 1934: 61 ( Philonthus ; illustration of aedeagus).
— Scheerpeltz, 1935a: 649 ( Philonthus ; Australia).
— Cameron, 1936: 6 ( Philonthus ; Austral Islands).
— C. Koch, 1936a: 75 ( Philonthus ; Libya).
— Wüsthoff, 1937a: 347 ( Philonthus ; illustration of aedeagus).
— Tottenham, 1937a: 190 ( Philonthus ; specimens of longicornis in Stephens’ collection with
type labels are not longicornis ).
— Gridelli, 1939: 317 ( Philonthus ; Abyssinia).
— Bernhauer, 1940c: 4 ( Philonthus ; Azores).
— Méquignon, 1942a: 20 ( Philonthus ; Azores).
— Blackwelder, 1943: 410 ( Philonthus ; characters; Haiti; Puerto Rico; St. Kitts; Antigua;
Montserrat; Grenada; Argentina; Chile).
— Cameron, 1949b: 468 ( Philonthus ; Japan).
— Cameron, 1950b: 214 ( Philonthus ; Sudan).
— V. Hansen, 1952: 67 ( Philonthus ; subgenus Philonthus ; characters; Denmark).
— Smetana, 1953: 118 ( Philonthus ; Turkey).
— Coiffait, 1954: 165 ( Philonthus ; Canary Islands).
— Tottenham, 1955: 163 ( Philonthus ; subgenus Spatulonthus ; notes; examined no African
specimens).
— Scheerpeltz, 1956a: 484 ( Philonthus ; Belgian Congo).
— Scheerpeltz, 1956c: 1100 ( Philonthus ; Cyprus).
— Scheerpeltz, 1957: 154 ( Philonthus ; Tanganyika).
— Smetana, 1958: 189 ( Philonthus ; subgenus Philonthus ; characters; ecology; Czechoslovakia).
— Smetana, 1959: 159 ( Philonthus ; characters).
— Nakane, 1963a: 91 ( Philonthus ; characters; habitus photograph; Japan).
— Smetana, 1963b: 36 ( Philonthus ; subgenus Philonthus ; Madeira).
— Palm, 1963b: 86 ( Philonthus ; characters; Sweden; Denmark; Norway; Finland).
— Lohse, 1964: 183 ( Philonthus ; subgenus Philonthus ; characters; central Europe).
— Focarile, 1964: 59 ( Philonthus ; Italy).
— Horion, 1965: 158 ( Philonthus ; all Europe).
— Raianu, 1967: 268 ( Philonthus ; characters; ecological notes; Romania).
— Szujecki, 1968a: 730 ( Philonthus ; Poland).
— Smetana, 1970: 59 ( Philonthus ; distributional summary; Azores).
— Scheerpeltz, 1971c: 170 ( Philonthus ; Cameroon).
— Fagel, 1972: 103 ( Spatulonthus ; notes; Sainte Helene).
— Moore and Legner, 1974a: 39 ( Philonthus ; spread by commerce).
— Coiffait, 1974: 311 ( Spatulonthus ; characters; Europe; Britain; Scandinavia; Anatolia).
— Scheerpeltz, 1974: 146 ( Philonthus ; Basutoland; South Africa).
— Scheerpeltz, 1974c: 22 ( Philonthus ; Ethiopia; Sudan).
— Moore, 1975c: 106 ( Philonthus ; Hawaii).
— Coiffait, 1976: 217 ( Spatulonthus ; Tahiti).
— Pace and Zanetti, 1977: 299 ( Spatulonthus ; Italy).
— Pope, 1977: 29 ( Philonthus ; Britain).
— Marcuzzi, 1977: 12 ( Philonthus ; Venezuela).
— Frank, 1982: 41 ( Philonthus ; list of parasites).
— Coiffait, 1984e: 374 ( Spatulonthus ; Nepal).
— Tóth, 1984: 68 ( Philonthus ; characters; Hungary).
— Yuh, Paik, Kwon, and Lee, 1985: 241 ( Philonthus ; Korea).
— Coiffait, 1986: 394 ( Spatulonthus ; Saudi Arabia).
— Last, 1987: 31 ( Philonthus ; Papua New Guinea).
— Lucht, 1987: 106 ( Philonthus ; checklist; Germany; Poland; Czechoslovakia; Austria;
Switzerland; France; Benelux; Denmark; Sweden).
— Borges, 1990: Table II ( Spatulonthus ; Azores).
— Kuschel, 1990: 26 ( Philonthus ; introduced, Palaearctic species; winged; habitat; New Zealand).

— Lecoq, 1990: 203 ( Spatulonthus ; Mascarene Islands). — Barrion, Litsinger, and Last, 1991: 1078 ( Philonthus ; associated with rice agroecosystem). — Pilon and Zanetti, 1991: 64 ( Philonthus ; Italy ). — J. Li, 1993: 37 ( Philonthus ; China ). — Smetana, 1993b: 48 ( Philonthus ; subgenus Spatulonthus ; Czech Republic ; Slovakia ). — Nishida, 1994: 73 ( Philonthus ; Hawaii ). — Hernández, Outerelo, and Gamarra, 1994: 188 ( Spatulonthus ; checklist of species; Canary Islands ). — Yadav, Chand, and Yadav, 1995: 6 ( Philonthus ; morphology of mitotic and meiotic chromosomes). — Ciceroni and Zanetti, 1995: 28 ( Philonthus ; Italy ). — Smetana, 1995e: 244 ( Philonthus ; characters; Manitoba; Nova Scotia; Ontario; Quebec; Alabama; Alaska; Arizona; Arkansas; California; Colorado; Florida; Georgia; Illinois; Indiana; Kansas; Maine; Maryland; Massachusetts; Michigan; Missouri; Nevada; New Jersey; New Mexico ; New York; North Carolina; Oregon; Pennsylvania; South Carolina; Texas; Utah; Vermont; Virginia; West Virginia; Washington; Wisconsin; Mexico ). — Hu and Frank, 1995b: 584 ( Philonthus ; surface structure of eggs). — Desender and Baert, 1996: 43 ( Philonthus ; Easter Island). — Hu and Frank, 1997 ( Philonthus ; predation of eggs and larvae of Haematobia irritans ; Florida ). — Nishida, 1997: 62 ( Philonthus ; Hawaii ). — Smithers, 1998: 16 ( Philonthus ; references; Norfolk Island ). — Kashcheev, 1999a: 57 ( Philonthus ; Kazakhstan ). — Staniec, 1999a: 56 ( Philonthus ; characters of pupa; Poland ). - DISTRIBUTION : Cosmopolitan. scybalarius Nordmann, 1837: 94 ( Philonthus ; Type locality: Berolinum ). — Erichson, 1839a: 465 ( Philonthus ; synonym of opacus ). — P. Lucas, 1846: 112 ( Philonthus ; synonym of varians ). — Wollaston, 1857: 189 ( Philonthus ; habitat; Madeira Islands ). — Guérin-Méneville, 1857: 40 ( Philonthus ; synonym of varians ). — Kraatz, 1857d: 601 ( Philonthus ; characters; Germany ). — Redtenbacher, 1857: 985 ( Philonthus ; characters; Austria ). — Kraatz, 1859: 86 ( Philonthus ; characters; Ceylon ). — Thomson, 1860: 164 ( Philonthus ; characters; Scandinavia). — Wollaston, 1864: 571 ( Philonthus ; notes; Canary Islands ). — Wollaston, 1865: 492 ( Philonthus ; habitat; notes; Madeira Islands ; Canary Islands ). — Crotch, 1867a: 383 ( Philonthus ; Azores ). — Wollaston, 1867: 237 ( Philonthus ; characters; notes; Cape Verde Islands). — Gemminger and Harold, 1868: 590 ( Philonthus ; synonym of longicornis ). — Wollaston, 1871: 302 ( Philonthus ; synonym of longicornis ). — Fauvel, 1874: 254 [= 1874a: 480] ( Philonthus ; synonym of longicornis ). — Sharp, 1874: 44 ( Philonthus ; notes; Japan ). — Redtenbacher, 1874: 213 ( Philonthus ; characters; Austria ). — Seidlitz, 1875: 277 ( Philonthus ; characters; Baltic region). — Mulsant and Rey, 1876: 472 [= 1877: 328] ( Philonthus ; characters; France ). — Fauvel, 1877: 264 ( Philonthus ; synonym of longicornis ). — Sharp, 1885: 416 ( Philonthus ; synonym of longicornis ). — Blackburn and Sharp, 1885: 222 ( Philonthus ; habitat; Hawaii ). — Ganglbauer, 1895: 454 ( Philonthus ; synonym of longicornis ). — Fauvel, 1902b: 112 ( Philonthus ; synonym of longicornis ). — Fauvel, 1903: 159 ( Philonthus ; synonym of longicornis ). — Fauvel, 1904a: 59 ( Philonthus ; synonym of longicornis ). — Sharp, 1908: 547 ( Philonthus ; Hawaii ). — Bernhauer and Schubert, 1914: 345 ( Philonthus ; synonym of longicornis ). — Cameron, 1932: 127 ( Philonthus ; synonym of longicornis ). — Scheerpeltz, 1933: 1349 ( Philonthus ; synonym of longicornis ). — Blackwelder, 1943: 410 ( Philonthus ; synonym of longicornis ).