<document id="8037DDA0A1217FF69152E93643EC374B" ID-CLB-Dataset="310020" ID-DOI="10.20362/am.015006" ID-GBIF-Dataset="87c63777-a328-4406-95b9-cae9677ba89e" ID-ISSN="2462-2362" ID-Zenodo-Dep="15473703" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1747749837623" checkinUser="felipe" docAuthor="Hamer, Matthew T., Lee, Jonathan Hon Chung, Tse, Cheung Yau Leo, Silva, Thiago S. R. &amp; Guénard, Benoit" docDate="2022" docId="03B687833139FFC9FF677AE3FDB5FB17" docLanguage="en" docName="AsianMyrmecol.15.e015006.pdf" docOrigin="Asian Myrmecology (e 015006) 15" docSource="http://dx.doi.org/10.5281/zenodo.11212378" docStyle="DocumentStyle:5808DD0968C355DB853299CB36ADE618.1:AsianMyrmecol.2019-.journal_article" docStyleId="5808DD0968C355DB853299CB36ADE618" docStyleName="AsianMyrmecol.2019-.journal_article" docStyleVersion="1" docTitle="Tetramorium insolens" docType="treatment" docVersion="9" lastPageNumber="109" masterDocId="FF8FFFFB3155FFA5FFA77A23FFAFFFEF" masterDocTitle="Remarkable diversity in a little red dot: a comprehensive checklist of known ant species in Singapore (Hymenoptera: Formicidae) with notes on ecology and taxonomy" masterLastPageNumber="152" masterPageNumber="1" pageNumber="109" updateTime="1748294324377" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BC5C9F4EE9DBD36F728625E51FCBCF24">Remarkable diversity in a little red dot: a comprehensive checklist of known ant species in Singapore (Hymenoptera: Formicidae) with notes on ecology and taxonomy</mods:title>
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<mods:namePart id="FCA4B02BA78ABE981751B84A91D71654">Hamer, Matthew T.</mods:namePart>
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<mods:namePart id="5028BFA4A649F8671510FF0E1B8E23F5">Lee, Jonathan Hon Chung</mods:namePart>
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<mods:namePart id="B3AC3C9D7247E718FFEC761D3CCF4681">Tse, Cheung Yau Leo</mods:namePart>
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<mods:namePart id="944BB3D3937E2A5E2E25592D7139BF21">Silva, Thiago S. R.</mods:namePart>
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<treatment id="03B687833139FFC9FF677AE3FDB5FB17" ID-DOI="http://doi.org/10.5281/zenodo.15474469" ID-GBIF-Taxon="262320332" ID-Zenodo-Dep="15474469" LSID="urn:lsid:plazi:treatment:03B687833139FFC9FF677AE3FDB5FB17" httpUri="http://treatment.plazi.org/id/03B687833139FFC9FF677AE3FDB5FB17" lastPageNumber="109" pageId="108" pageNumber="109" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="C305651E3139FFC9FF677AE3FDFEFF37" box="[192,593,191,216]" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="8BA036953139FFC9FF677AE3FDFEFF37" blockId="108.[192,593,191,216]" box="[192,593,191,216]" pageId="108" pageNumber="109">
<heading id="D0E881F93139FFC9FF677AE3FDFEFF37" box="[192,593,191,216]" fontSize="10" level="2" pageId="108" pageNumber="109" reason="4">
<taxonomicName id="4C1F4D163139FFC9FF677AE3FDFEFF37" ID-CoL="55PNJ" authority="(Smith, 1861)" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[192,593,191,216]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B96BEA873139FFC9FF677AE3FDFEFF37" bold="true" box="[192,593,191,216]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677AE3FE1EFF38" bold="true" box="[192,433,191,216]" italics="true" pageId="108" pageNumber="109">Tetramorium insolens</emphasis>
(
<bibRefCitation id="EF8E4B643139FFC9FE187AE3FDE6FF37" author="Smith F" box="[447,585,192,216]" pageId="108" pageNumber="109" pagination="36 - 48" refId="ref110632" refString="Smith F, 1861. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Ceram, Celebes, Ternate, and Gilolo. [part]. Journal and Proceedings of the Linnean Society of London. Zoology 6: 36-48." type="journal article" year="1861">Smith, 1861</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C305651E3139FFC9FF677B23FDEBFD18" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="8BA036953139FFC9FF677B23FD2CFDF7" blockId="108.[192,719,256,1272]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677B23FE0CFEF7" bold="true" box="[192,419,256,280]" pageId="108" pageNumber="109">Material examined.</emphasis>
<materialsCitation id="3B773CC83139FFC9FE0C7B23FD2AFED7" ID-GBIF-Occurrence="5163310607" collectingDate="2016-11-01" collectionCode="ZRC" collectorName="G. W. Yong" location="Kranji Road" pageId="108" pageNumber="109" specimenCode="ENT00048415" specimenCount="1">
<location id="8EC0604E3139FFC9FE0C7B23FD9BFEF7" LSID="urn:lsid:plazi:treatment:03B687833139FFC9FF677AE3FDB5FB17:8EC0604E3139FFC9FE0C7B23FD9BFEF7" box="[427,564,256,280]" name="Kranji Road" pageId="108" pageNumber="109">Kranji Road</location>
, 
<date id="FFA110553139FFC9FDE57B23FD68FEF7" box="[578,711,256,280]" pageId="108" pageNumber="109" value="2016-11-01">
<collectingDate id="EFE5E9BD3139FFC9FDE57B23FD68FEF7" box="[578,711,256,280]" pageId="108" pageNumber="109" value="2016-11-01">1 Nov 2016</collectingDate>
</date>
, 
<collectorName id="26EA53433139FFC9FF577B03FEC9FED7" box="[240,358,288,312]" pageId="108" pageNumber="109">G.W. Yong</collectorName>
leg., 
<collectionCode id="ED0EAE503139FFC9FE067B03FE79FED7" box="[417,470,288,312]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ZRC</collectionCode>
_ 
<specimenCode id="DBB99EEE3139FFC9FE437B03FD2AFED7" box="[484,645,288,312]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ENT00048415</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B773CC83139FFC9FD287B03FDF1FE57" ID-GBIF-Occurrence="5163310157" collectingDate="2018-03-14" collectionCode="ZRC" collectorName="M. S. Foo &amp; W. Wang" country="Singapore" location="Singapore" pageId="108" pageNumber="109" specimenCode="ENT00000907" specimenCount="1">
alates and workers, National University of 
<collectingCountry id="F30876053139FFC9FDC47B63FD62FEB7" box="[611,717,320,344]" name="Singapore" pageId="108" pageNumber="109">Singapore</collectingCountry>
campus (Clementi/Kent Ridge, multiple sites), Apr-Aug 2015, 
<collectorName id="26EA53433139FFC9FE077BA3FDA7FE78" box="[416,520,384,408]" pageId="108" pageNumber="109">M.S. Foo</collectorName>
&amp; 
<collectorName id="26EA53433139FFC9FD887BA3FD3BFE77" box="[559,660,384,408]" pageId="108" pageNumber="109">W. Wang</collectorName>
leg., 
<collectingMethod id="525E4E823139FFC9FF577B83FEC0FE58" box="[240,367,416,440]" pageId="108" pageNumber="109">malaise trap</collectingMethod>
, 
<collectionCode id="ED0EAE503139FFC9FEDF7B83FE04FE57" box="[376,427,416,440]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ZRC</collectionCode>
_BDP (multiple);
</materialsCitation>
<materialsCitation id="3B773CC83139FFC9FDC47B83FDD0FDF7" ID-GBIF-Occurrence="5163309526" collectingDate="2018-03-14" collectionCode="ZRC" collectorName="M. S. Foo &amp; W. Wang" country="Singapore" location="Singapore" pageId="108" pageNumber="109" specimenCode="ENT00000907" specimenCount="1">
queen and workers, 
<location id="8EC0604E3139FFC9FEF37BE3FD09FE38" LSID="urn:lsid:plazi:treatment:03B687833139FFC9FF677AE3FDB5FB17:8EC0604E3139FFC9FEF37BE3FD09FE38" box="[340,678,448,472]" country="Singapore" name="Sungei Buloh Wetland Reserve" pageId="108" pageNumber="109">Sungei Buloh Wetland Reserve</location>
, 
<date id="FFA110553139FFC9FD167BE3FEF3FE17" pageId="108" pageNumber="109" value="2018-03-14">
<collectingDate id="EFE5E9BD3139FFC9FD167BE3FEF3FE17" pageId="108" pageNumber="109" value="2018-03-14">14 Mar 2018</collectingDate>
</date>
, 
<collectorName id="26EA53433139FFC9FECF7BC3FE64FE17" box="[360,459,480,504]" pageId="108" pageNumber="109">W. Wang</collectorName>
&amp; 
<collectorName id="26EA53433139FFC9FE487BC3FDFBFE18" box="[495,596,480,504]" pageId="108" pageNumber="109">M.S. Foo</collectorName>
leg., WW-SG18- 
<collectorName id="26EA53433139FFC9FE937823FE26FDF7" box="[308,393,512,536]" pageId="108" pageNumber="109">Tetram</collectorName>
1, 
<collectionCode id="ED0EAE503139FFC9FE3C7823FE7FFDF7" box="[411,464,512,536]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ZRC</collectionCode>
_ 
<specimenCode id="DBB99EEE3139FFC9FE797823FDD0FDF7" box="[478,639,512,536]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ENT00000907</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA036953139FFC9FF677803FD15FDD7" blockId="108.[192,719,256,1272]" box="[192,698,544,568]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677803FDECFDD7" bold="true" box="[192,579,544,568]" pageId="108" pageNumber="109">Material not physically examined.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA036953139FFC9FF677863FDA5FD77" blockId="108.[192,719,256,1272]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677863FE92FDB7" bold="true" box="[192,317,576,600]" pageId="108" pageNumber="109">Literature.</emphasis>
<bibRefCitation id="EF8E4B643139FFC9FEE47863FDB9FDB7" author="Wang WYL &amp; Srivathsan A &amp; Foo M &amp; Yamane SK &amp; Meier R" box="[323,534,576,600]" pageId="108" pageNumber="109" pagination="490 - 501" refId="ref111460" refString="Wang WYL, Srivathsan A, Foo M, Yamane SK and Meier R, 2018 a. Sorting specimen-rich invertebrate samples with cost-effective NGS barcodes: validating a reverse workflow for specimen processing. Molecular Ecology Resources 18 (3): 490-501." type="journal article" year="2018">Wang et al. (2018a)</bibRefCitation>
[as 
<taxonomicName id="4C1F4D163139FFC9FDE57862FEC6FD97" baseAuthorityName="Smith" baseAuthorityYear="1861" class="Insecta" family="Formicidae" genus="Tetramorium" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B96BEA873139FFC9FDE57862FD63FDB7" box="[578,716,577,600]" italics="true" pageId="108" pageNumber="109">Tetramorium</emphasis>
cf. 
<emphasis id="B96BEA873139FFC9FEB57843FEC6FD97" box="[274,361,608,632]" italics="true" pageId="108" pageNumber="109">insolens</emphasis>
</taxonomicName>
– 
<collectionCode id="ED0EAE503139FFC9FE237843FE17FD97" box="[388,440,608,632]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="108" pageNumber="109">ZRC</collectionCode>
material determined as 
<taxonomicName id="4C1F4D163139FFC9FD1E7842FEE8FD77" baseAuthorityName="Smith" baseAuthorityYear="1861" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B96BEA873139FFC9FD1E7842FEE8FD77" italics="true" pageId="108" pageNumber="109">T. insolens</emphasis>
</taxonomicName>
post publication].
</paragraph>
<paragraph id="8BA036953139FFC9FF677883FDEBFD18" blockId="108.[192,719,256,1272]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677883FE99FD57" bold="true" box="[192,310,672,696]" pageId="108" pageNumber="109">Localities.</emphasis>
Kranji Road; National University of 
<collectingCountry id="F30876053139FFC9FF5778E3FEF0FD37" box="[240,351,704,728]" name="Singapore" pageId="108" pageNumber="109">Singapore</collectingCountry>
campus (Clementi/Kent Ridge); Sungei Buloh Wetland Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C305651E3139FFC9FF677923FD12FC18" pageId="108" pageNumber="109" type="biology_ecology">
<paragraph id="8BA036953139FFC9FF677923FD12FC18" blockId="108.[192,719,256,1272]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677923FED3FCF7" bold="true" box="[192,380,768,792]" pageId="108" pageNumber="109">Habitat/Ecology.</emphasis>
<taxonomicName id="4C1F4D163139FFC9FE257922FDC8FCF7" baseAuthorityName="Smith" baseAuthorityYear="1861" box="[386,615,768,792]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B96BEA873139FFC9FE257922FDC8FCF7" box="[386,615,768,792]" italics="true" pageId="108" pageNumber="109">Tetramorium insolens</emphasis>
</taxonomicName>
is associated with a wide variety of different habitats in 
<collectingCountry id="F30876053139FFC9FEAA7963FED3FCB7" box="[269,380,832,856]" name="Singapore" pageId="108" pageNumber="109">Singapore</collectingCountry>
, from mangrove back forest to urban cultivated patches, also disturbed secondary forest fragments in urban or semi-urban settings. In back mangroves, the ants have been found in a thin carton nest on the underside of a thick branch of a living tree.
</paragraph>
</subSubSection>
<subSubSection id="C305651E3139FFC9FF677E23FDB5FB17" pageId="108" pageNumber="109" type="discussion">
<paragraph id="8BA036953139FFC9FF677E23FDB5FB17" blockId="108.[192,719,256,1272]" pageId="108" pageNumber="109">
<emphasis id="B96BEA873139FFC9FF677E23FE80FBF7" bold="true" box="[192,303,1024,1048]" pageId="108" pageNumber="109">Remarks.</emphasis>
This species is considered a relatively widespread ‘pan-global’ tramp species originating from 
<collectingCountry id="F30876053139FFC9FEDA7E63FE47FBB7" box="[381,488,1088,1112]" name="Indonesia" pageId="108" pageNumber="109">Indonesia</collectingCountry>
and probably introduced elsewhere (
<bibRefCitation id="EF8E4B643139FFC9FE0C7E43FD9BFB97" author="Bolton B" box="[427,564,1120,1144]" pageId="108" pageNumber="109" pagination="67 - 151" refId="ref103377" refString="Bolton B, 1977. The ant tribe Tetramoriini (Hymenoptera: Formicidae). The genus Tetramorium Mayr in the Oriental and Indo-Australian regions, and in Australia. Bulletin of the British Museum (Natural History). Entomology 36: 67-151." type="journal article" year="1977">Bolton 1977</bibRefCitation>
). It is morphologically similar to and often misidentified as 
<taxonomicName id="4C1F4D163139FFC9FEA87E82FE03FB57" baseAuthorityName="Nylander" baseAuthorityYear="1846" box="[271,428,1184,1208]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="bicarinatum">
<emphasis id="B96BEA873139FFC9FEA87E82FE03FB57" box="[271,428,1184,1208]" italics="true" pageId="108" pageNumber="109">T. bicarinatum</emphasis>
</taxonomicName>
, but may be differentiated by characters detailed in ‘Remarks’ under the latter species section above.
</paragraph>
</subSubSection>
</treatment>
</document>