<document ID-CLB-Dataset="6051" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62615" ID-GBIF-Dataset="2a9d2591-e034-4433-8be1-517c6921c937" ID-PMC="PMC9425446" ID-Pensoft-Pub="1313-2970-1044-449" ID-Pensoft-UUID="9FB3984FE63050AEBDD824E47EF5EF42" ID-PubMed="36187209" ID-ZooBank="E2CB7962B7904B95BD003650F11AE2D" ModsDocID="1313-2970-1044-449" checkinTime="1623875390689" checkinUser="pensoft" docAuthor="Assmann, Thorsten, Boutaud, Esteve, Buse, Joern, Drees, Claudia, Friedman, Ariel-Leib-Leonid, Harry, Ingmar, Khoury, Fares, Orbach, Eylon, Renan, Ittai, Schmidt, Constantin, Schmidt, Kilian, Wrase, David W. &amp; Zumstein, Pascale" docDate="2021" docId="03964FA4A59E5E6EBEBCD57B19835F74" docLanguage="en" docName="ZooKeys 1044: 449-478" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62615" docTitle="Agonum monachum subsp. syriacum Schmidt &amp; Liebherr 2009" docType="treatment" docVersion="6" id="9FB3984FE63050AEBDD824E47EF5EF42" lastPageNumber="449" masterDocId="9FB3984FE63050AEBDD824E47EF5EF42" masterDocTitle="The ground beetle tribe Platynini Bonelli, 1810 (Coleoptera, Carabidae) in the southern Levant: dichotomous and interactive identification tools, ecological traits, and distribution" masterLastPageNumber="478" masterPageNumber="449" pageNumber="449" updateTime="1732492657825" updateUser="ExternalLinkService">
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<mods:title id="4EF3E359B430E6A4DB5E3BFFBB9974DA">The ground beetle tribe Platynini Bonelli, 1810 (Coleoptera, Carabidae) in the southern Levant: dichotomous and interactive identification tools, ecological traits, and distribution</mods:title>
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<mods:namePart id="AD49130C8BE508C2250C418E58CC8378">Assmann, Thorsten</mods:namePart>
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<mods:affiliation id="46EEAA14C635D5DE0DBE957A0166DC4C">Ecosystem Monitoring, Research and Wildlife Conservation (SB 23 Invertebrates and Biodiversity), Black Forest National Park, Kniebisstrasse 67, D- 72250 Freudenstadt, Germany</mods:affiliation>
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<mods:affiliation id="DC47339D398631B49626D8ECEB2340ED">School of Life Sciences, University of Sussex, Brighton, BN 1 9 QG, United Kingdom</mods:affiliation>
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<mods:namePart id="82824B00D5CF46078A63A0926179F086">Khoury, Fares</mods:namePart>
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<mods:namePart id="D2D54A71CA535984A1D829CA64E14BD4">Orbach, Eylon</mods:namePart>
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<mods:affiliation id="3BA0B436962A90597787D5FCAD2B0689">Steinhardt Museum of Natural History, Tel Aviv University, Klauzner 12, Tel Aviv, IL- 69978, Israel</mods:affiliation>
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<mods:namePart id="36E24B170BBF39B2190BAC36F1A556C1">Schmidt, Constantin</mods:namePart>
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<mods:namePart id="FAF5C2F110FA0E59F5666FC40B19A89D">Schmidt, Kilian</mods:namePart>
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<mods:namePart id="F5CD507009A1E40EDA987E949D7A324B">Wrase, David W.</mods:namePart>
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<mods:namePart id="3C67CB2765AD4F2C2247CFD13BA00E1B">Zumstein, Pascale</mods:namePart>
<mods:affiliation id="5637139179B5A48C3454420C84D29809">Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21335 Lueneburg, Germany</mods:affiliation>
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<treatment id="03964FA4A59E5E6EBEBCD57B19835F74" ID-GBIF-Taxon="182754001" LSID="urn:lsid:plazi:treatment:03964FA4A59E5E6EBEBCD57B19835F74" httpUri="http://treatment.plazi.org/id/03964FA4A59E5E6EBEBCD57B19835F74" lastPageNumber="449" pageId="0" pageNumber="449" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Platynini">
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<paragraph id="AEC03AB992C6A96002269E12349F315B" pageId="0" pageNumber="449">
<taxonomicName id="A03D69948A50DB16C0AA0930811E92FF" ID-CoL="8MH89" LSID="03964FA4-A59E-5E6E-BEBC-D57B19835F74" authority="Schmidt &amp; Liebherr, 2009" authorityName="Schmidt &amp; Liebherr" authorityYear="2009" class="Insecta" family="Carabidae" genus="Agonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonum monachum subsp. syriacum" order="Coleoptera" pageId="0" pageNumber="449" phylum="Arthropoda" rank="subSpecies" species="monachum" subSpecies="syriacum">Agonum (s. str.) monachum syriacum Schmidt &amp; Liebherr, 2009</taxonomicName>
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<paragraph id="6C340C8B190DE0B4CEE2FCCA821A24F4" pageId="0" pageNumber="449">Dispersal power.</paragraph>
<paragraph id="B87DAADE7189C5D56BA6C5EDB8A9D30B" pageId="0" pageNumber="449">Fully winged and flight-active (pers. obs.).</paragraph>
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<subSubSection id="E38BEBA0836A2F5421D7F0884390B03E" pageId="0" pageNumber="449" type="habitat">
<paragraph id="3C7462047B39C7B316BEBDEA8A108134" pageId="0" pageNumber="449">Habitat.</paragraph>
<paragraph id="85FE5CB9103CB787A63E996C453176AA" pageId="0" pageNumber="449">
Known only from winter ponds (Fig. 
<figureCitation id="4067E743B6E94BBD5FC21F9AFC651169" captionStart="Figures 31–34" captionStartId="F10" captionText="Figures 31 - 34. Habitats of Platynini species in the southern Levant 31 Merom Golan (Golan Heights): water reservoir, habitat of Agonum rugicolle, A. sordidum, A. marginatum, A. viridicupreum, A. nigrum, and Anchomenus dorsalis infuscatus 32 Ein Akev (Central Negev): headwater stream in desert, habitat of Agonum nigrum 33 Breikhat Ya'ar (Coastal Plain, east of Hadera): winter pond, habitat of Agonum monachum syriacum, A. viridicupreum, A. nigrum, and Anchomenus dorsalis infuscatus 34 Nahal Bezet (Upper Galilee): softwood floodplain woodland with Oleander (Nerium oleander), habitat of Atranus ruficollis." figureDoi="10.3897/zookeys.1044.62615.figures31-34" httpUri="https://binary.pensoft.net/fig/554582" pageId="0" pageNumber="449">31</figureCitation>
) and intact large wetlands (e.g., swamps north of the Sea of Galilee).
</paragraph>
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<paragraph id="16321384C4FF87D29C95A1324A36616E" pageId="0" pageNumber="449">Phenology.</paragraph>
<paragraph id="6A856FBC8B365F6EAD16577F887E2A9E" pageId="0" pageNumber="449">Spring breeder, adults hibernate (pers. obs.).</paragraph>
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<subSubSection id="F3E689B9568D4257DED494C289AFD4EE" pageId="0" pageNumber="449" type="distribution">
<paragraph id="B13A1335DDC544C645A58E1249A867E2" pageId="0" pageNumber="449">Distribution range.</paragraph>
<paragraph id="A2D7D55C71E871F8CD873A175DC5A867" pageId="0" pageNumber="449">
The species is distributed from Southwest Europe to Central Asia, southwards to Israel and Iran (
<bibRefCitation id="FEA78815D2FF5FC9679E13967E912ADA" author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt 2017</bibRefCitation>
). The subspecies 
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is known from Southeast Turkey to Israel (
<bibRefCitation id="312AA588F64FC6B697DA3454BC225121" author="Schmidt, J" journalOrPublisher="VERNATE" pageId="0" pageNumber="449" pagination="225 - 257" refId="B72" refString="Schmidt, J, Liebherr, JK, 2009. Beitraege zur Systematik und Verbreitung palaearktischer Arten der Platynini (Insecta: Coleoptera: Carabidae). VERNATE 28: 225 - 257" title="Beitraege zur Systematik und Verbreitung palaearktischer Arten der Platynini (Insecta: Coleoptera: Carabidae)." volume="28" year="2009">Schmidt and Liebherr 2009</bibRefCitation>
) and from Iran (
<bibRefCitation id="96285E691F8979E9649699A3AC8A28B1" author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3E071E97E01CB101DA597778E5DEE3B8" pageId="0" pageNumber="449" type="distribution">
<paragraph id="6930F1539A0EB25F7E64208541B67454" pageId="0" pageNumber="449">Distribution in the southern Levant.</paragraph>
<paragraph id="BE234553F5C646211C52558410E28AB9" pageId="0" pageNumber="449">
In the southern Levant recorded exclusively from four sites in Israel, three of them from winter pools in the Central Coastal Plain (Berekhat 
<normalizedToken id="48DCB094934C3930CD5D5A788965ACA7" originalValue="Ya’ar">Ya'ar</normalizedToken>
(sometimes also written Breikhat 
<normalizedToken id="02273C25B919B83C7CF62E38B32DEF4B" originalValue="Ya’ar">Ya'ar</normalizedToken>
), Dora Pool and 
<normalizedToken id="11766B18A4313830077BC1F32B6A9137" originalValue="Ga’ash">Ga'ash</normalizedToken>
Pool; from the latter two sites only single specimens). A large population exists also in the swamps north of the Sea of Galilee.
</paragraph>
</subSubSection>
<subSubSection id="1FD1B5C44231BEC460DB061EB6AFF816" pageId="0" pageNumber="449" type="taxonomic notes">
<paragraph id="E03FA9871609313A75D70E6B9C53F260" pageId="0" pageNumber="449">Taxonomic notes.</paragraph>
<paragraph id="6DCFF6CB2F53C91B1FC4A6D0F8AB0648" pageId="0" pageNumber="449">
<bibRefCitation id="5F9EA2495E5650DFC789792F7CBF23E3" author="Schmidt, J" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="449" pagination="642 - 675" refId="B71" refString="Schmidt, J, 2017. Tribe Platynini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga). Brill, Leiden, Boston: 642 - 675" title="Tribe Platynini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Aechostemata - Myxophaga - Adephaga)." year="2017">Schmidt (2017)</bibRefCitation>
listed the nominate subspecies from Israel, but we know only the subspecies subspecies 
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from the southern Levant. All specimens we know from the southern Levant show a greenish metallic luster on the upper side, despite the indication in the original description.
</paragraph>
</subSubSection>
<subSubSection id="9AC2CFAD14E30D3A8713674587807C7F" pageId="0" pageNumber="449" type="conservation">
<paragraph id="588AC165244FE3DF60F35ADBA105F0C2" pageId="0" pageNumber="449">Conservation.</paragraph>
<paragraph id="B4A9E97E5B0E590596DEDCCAC0E5BA71" pageId="0" pageNumber="449">
In contrast to 
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<emphasis id="633378DF1F937BA196EF719561EDE94E" italics="true" pageId="0" pageNumber="449">A. nigrum</emphasis>
</taxonomicName>
a rarely recorded species; we know only two larger populations (Sea of Galilee, Berekhat 
<normalizedToken id="B0DA66F6DFCE734230538ACC7A559F25" originalValue="Ya’ar">Ya'ar</normalizedToken>
). Threatened by habitat loss, drainage, and water table drawdown.
</paragraph>
</subSubSection>
</treatment>
</document>