Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae) Author WANG, YA-LIAN Author LI, BING-XUE Author CUI, MENG-DI Author ZHANG, YUAN Author WANG, LE Author ZHANG, CAI-HONG Author TSAUR, SHUN-CHERN Author CHEN, HONG-WEI Author HUANG, JIA text Zootaxa 2023 2023-03-07 5250 1 1 109 http://dx.doi.org/10.11646/zootaxa.5250.1.1 journal article 53812 10.11646/zootaxa.5250.1.1 da8acfed-df61-403e-9a49-7f856ce56196 1175­5334 7705657 4729EAF3-8B78-4AAD-B373-9179D008F239 Stegana ( Steganina ) marenubila Cui & Chen , sp. nov. ( Figs. 5A, 5B , 21 ) Diagnosis. This species resembles S . ( S .) crinata Zhang & Chen, 2015 in the shape of epandrium, surstylus, and cercus (as fig. 2 in Zhang & Chen 2015 , Fig. 21A ), but can be distinguished from the latter by having the aedeagus with 3 pair of basally fused processes: ventral pair small, medial pair slender, and dorsal pair strong ( Fig. 21B, C ). FIGURE 21. Stegana ( Steganina ) marenubila Cui & Chen , sp. nov. , male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view. Description. Male and female: Frons yellow in male, brown in female. First flagellomere yellow in male, brown in female. Face black, with 1 yellow band medially; facial carina absent. Clypeus black. Palpus mostly yellow. Mesonotum and scutellum yellow in male ( Fig. 5A, B ), brownish in female. Katepisternum yellowish ( Fig. 5B). Halter mostly grey ( Fig. 5B ). Fore femur with 3 thin, long setae on distal part of posteroventral surface. Abdominal tergites yellow ( Fig. 5A, B ), 4th to 6th tergites brownish medially in female; sternites yellow, about 2 times as wide as long. Male terminalia ( Fig. 21 ): Epandrium with ca . 8 setae on dorsal to posterior portions per side ( Fig. 21A ). Surstylus lacking pubescence ( Fig. 21A ). Cercus with much pubescence ( Fig. 21A ). Hypandrium slightly expanded anteromedially ( Fig. 21B, C ). Pregonite blurry or absent ( Fig. 21B, C ). Junction between hypandrium and aedeagus with 3 sensilla per side ( Fig. 21B, C ). Measurements and indices. BL = 2.45 mm in holotype ( 2.73 in 1♀ paratype ), ThL = 1.21 mm (1.19), WL = 2.56 mm (2.75), WW = 0.93 mm (1.12), arb = 7/4 (7/5), avd = 0.73 (0.67), adf = 1.07 (1.21), flw = 1.42 (1.69), FW/ HW = 0.32 (0.37), ch/o = 0.24 (0.24), prorb = 0.96 (0.90), rcorb = 1.20 (0.54), vb = 0.57 (0.46), dcl = 0.28 (1.22), presctl = 0.38 (0.38), sctl = damaged (2.63), sterno = 0.41 (0.46), orbito = 2.22 (1.89), dcp = 0.53 (0.30), sctlp = 1.50 (1.39), C = 2.20 (2.16), 4c = 1.62 (1.02), 4v = 2.49 (1.75), 5x = 1.29 (1.42), ac = 10.75 (8.46), M = 0.93 (0.56), C3F = 0.64 (0.60). Type materials. Holotype ♂ ( SCAU , No. 125411, DNA #2347), CHINA : Manghai , Mangshi , Yunnan , 24°07'03''N , 98°19'04''E , alt. 1720m , 8.xi.2017 , ex tree trunk, H.W. Chen . Paratype : 1♀ ( SCAU , No. 125412, DNA #2747), other data same as holotype. Distribution. China ( Yunnan ). Etymology. A combination of the Latin words " mare " (= sea) and " nubilus " (= nubilous), referring to the meaning of the type locality (Manghai: a sea of clouds).