<document id="25E1AFAF1CF0E68A07E160CCB77DE1E0" ID-DOI="10.5281/zenodo.5176513" ID-ISSN="1942-1354" ID-Zenodo-Dep="5176513" ID-ZooBank="068D2C0D-C92D-4E47-93FF-222CA4D5D824" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628329503398" checkinUser="felipe" docAuthor="Canepari, Claudio, Gordon, Robert D. &amp; Hanley, Guy A." docDate="2013" docId="038F8787FFB2FF9CFF7C5E96FBECFB20" docLanguage="en" docName="InsectaMundi.0312.1-91.pdf" docOrigin="Insecta Mundi 2013 (312)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Dilatitibialis tropicalis, new combination" docType="treatment" docVersion="4" lastPageNumber="29" masterDocId="FFB6FFFFFFADFFBCFFBC5B55FF8DFFFB" masterDocTitle="South American Coccinellidae (Coleoptera), Part XV: systematic revision of Dilatitibialis Duverger (Coccidulinae; Hyperaspidini)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="28" updateTime="1699044533877" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F1E926F5F51ED569AB219FCEEFFC8879">South American Coccinellidae (Coleoptera), Part XV: systematic revision of Dilatitibialis Duverger (Coccidulinae; Hyperaspidini)</mods:title>
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<mods:namePart id="2087D2F3A96B0A9A613458AADE355E2E">Canepari, Claudio</mods:namePart>
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<mods:namePart id="200B0124EC4845B70D0C6CA965A07816">Gordon, Robert D.</mods:namePart>
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<mods:title id="98E7D06214FC5954658EF41B830E542A">Insecta Mundi</mods:title>
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<subSubSection id="C33C651AFFB2FFA3FF7C5E96FC3DFA20" box="[192,944,1475,1499]" pageId="31" pageNumber="28" type="description">
<paragraph id="8B993691FFB2FFA3FF7C5E96FC3DFA20" blockId="31.[192,944,1475,1499]" box="[192,944,1475,1499]" pageId="31" pageNumber="28">
<heading id="D0D181FDFFB2FFA3FF7C5E96FC3DFA20" box="[192,944,1475,1499]" fontSize="10" level="2" pageId="31" pageNumber="28" reason="7">
18. 
<emphasis id="B952EA83FFB2FFA3FF565E96FC25FA20" bold="true" box="[234,936,1475,1499]" pageId="31" pageNumber="28">
<taxonomicName id="4C264D12FFB2FFA3FF565E96FD39FA20" authority="(Mulsant)" baseAuthorityName="Mulsant" box="[234,692,1475,1499]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="28" phylum="Arthropoda" rank="species" species="tropicalis" status="comb. nov.">
<emphasis id="B952EA83FFB2FFA3FF565E96FDAAFA20" bold="true" box="[234,551,1475,1499]" italics="true" pageId="31" pageNumber="28">Dilatitibialis tropicalis</emphasis>
(Mulsant)
</taxonomicName>
, 
<taxonomicNameLabel id="A26157F8FFB2FFA3FD035E96FC25FA20" box="[703,936,1475,1499]" pageId="31" pageNumber="28" rank="species">new combination</taxonomicNameLabel>
</emphasis>
.
</heading>
</paragraph>
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<subSubSection id="C33C651AFFB2FFA3FF7C5D56FB89F9C0" pageId="31" pageNumber="28" type="reference_group">
<paragraph id="8B993691FFB2FFA3FF7C5D56FD3FF9E0" blockId="31.[192,1028,1539,1595]" box="[192,690,1539,1563]" pageId="31" pageNumber="28">
<treatmentCitationGroup id="AB3611BFFFB2FFA3FF7C5D56FD3FF9E0" box="[192,690,1539,1563]" pageId="31" pageNumber="28">
<treatmentCitation id="0A871080FFB2FFA3FF7C5D56FD23F9E0" author="Mulsant, M. E." box="[192,686,1539,1563]" page="1038" pageId="31" pageNumber="28" year="1850">
<taxonomicName id="4C264D12FFB2FFA3FF7C5D56FD23F9E0" authority="Mulsant, 1850: 1038" authorityName="Mulsant, 1850:" authorityPageNumber="1038" authorityYear="1038" box="[192,686,1539,1563]" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="28" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B952EA83FFB2FFA3FF7C5D56FE20F9E0" box="[192,429,1539,1563]" italics="true" pageId="31" pageNumber="28">Cleothera tropicalis</emphasis>
<bibRefCitation id="EFB74B60FFB2FFA3FE085D56FD23F9E0" author="Mulsant, M. E." box="[436,686,1539,1563]" pageId="31" pageNumber="28" pagination="1 - 1104" refId="ref57076" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant, 1850: 1038</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B993691FFB2FFA3FF7C5D71FB89F9C0" blockId="31.[192,1028,1539,1595]" box="[192,1028,1571,1595]" pageId="31" pageNumber="28">
<treatmentCitationGroup id="AB3611BFFFB2FFA3FF7C5D71FB89F9C0" box="[192,1028,1571,1595]" pageId="31" pageNumber="28">
<treatmentCitation id="0A871080FFB2FFA3FF7C5D71FD6CF9C0" author="Korschefsky, R." box="[192,737,1571,1595]" page="198" pageId="31" pageNumber="28" year="1931">
<taxonomicName id="4C264D12FFB2FFA3FF7C5D71FD6CF9C0" authority=": Korschefsky 1931: 198" authorityName="Korschefsky" authorityPageNumber="198" authorityYear="1931" box="[192,737,1571,1595]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="28" phylum="Arthropoda" rank="species" species="tropicalis">
<emphasis id="B952EA83FFB2FFA3FF7C5D71FE4DF9C0" box="[192,448,1571,1595]" italics="true" pageId="31" pageNumber="28">Hyperaspis tropicalis</emphasis>
: 
<bibRefCitation id="EFB74B60FFB2FFA3FE725D76FD6CF9C0" author="Korschefsky, R." box="[462,737,1571,1595]" pageId="31" pageNumber="28" pagination="1 - 224" refId="ref56982" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118: 1 - 224." type="book chapter" year="1931">Korschefsky 1931: 198</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; 
<treatmentCitation id="0A871080FFB2FFA3FD505D76FB8DF9C0" author="Blackwelder, R. E." box="[748,1024,1571,1595]" page="448" pageId="31" pageNumber="28" year="1945">
<bibRefCitation id="EFB74B60FFB2FFA3FD505D76FB8DF9C0" author="Blackwelder, R. E." box="[748,1024,1571,1595]" pageId="31" pageNumber="28" pagination="343 - 550" refId="ref56508" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 448</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33C651AFFB2FF9CFF7C5D36FE5BFCC0" lastPageId="32" lastPageNumber="29" pageId="31" pageNumber="28" type="description">
<paragraph id="8B993691FFB2FF9CFF7C5D36FBC5FD20" blockId="31.[192,1441,1635,1916]" lastBlockId="32.[192,1441,195,763]" lastPageId="32" lastPageNumber="29" pageId="31" pageNumber="28">
<emphasis id="B952EA83FFB2FFA3FF7C5D36FEEEF980" bold="true" box="[192,355,1635,1659]" pageId="31" pageNumber="28">Description</emphasis>
. 
<emphasis id="B952EA83FFB2FFA3FECA5D36FE34F980" bold="true" box="[374,441,1635,1659]" pageId="31" pageNumber="28">Male</emphasis>
<typeStatus id="549D8833FFB2FFA3FE785D36FDBCF980" box="[452,561,1635,1659]" pageId="31" pageNumber="28" type="holotype">holotype</typeStatus>
. Length 
<quantity id="4CDE9B74FFB2FFA3FD1F5D36FC88F980" box="[675,773,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="31" pageNumber="28" unit="mm" value="3.1">3.1 mm</quantity>
, width 
<quantity id="4CDE9B74FFB2FFA3FCDB5D36FC44F980" box="[871,969,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="31" pageNumber="28" unit="mm" value="2.3">2.3 mm</quantity>
; body rounded, convex. Color yellow; pronotum with large, basomedian, dark brown macula, macula wide across base, narrowed anteriorly; elytron dark brown with 5 yellow spots arranged as in 
<figureCitation id="131D2A14FFB2FFA3FCE25DF6FC38F940" box="[862,949,1699,1723]" captionStart="Figures 86-101. 86-91" captionStartId="80.[192,286,1611,1633]" captionTargetBox="[207,1418,205,1583]" captionTargetId="figure-19@80.[192,1440,192,1618]" captionTargetPageId="80" captionText="Figures 86-101. 86-91) Dilatitibialis gravabilis. 86) Habitus. 87) Phallobase ventral. 88) Phallobase lateral. 89) Enlarged siphonal apex. 90) Sipho. 91) Female genitalia. 92-96) Dilatitibialis lillian. 92) Habitus. 93) Phallobase ventral. 94) Phallobase lateral. 95) Sipho. 96) Enlarged siphonal apex. 97-101) Dilatitibialis tropicalis. 97) Habitus. 98) Phallobase ventral. 99) Phallobase lateral. 100) Sipho. 101) Enlarged siphonal apex." figureDoi="http://doi.org/10.5281/zenodo.5176527" httpUri="https://zenodo.org/record/5176527/files/figure.png" pageId="31" pageNumber="28">Fig. 97</figureCitation>
; ventral surface with head, prosternum, meso- and metaventrites dark reddish yellow; abdomen dark yellowish brown medially, yellow laterally. Head punctures separated by about a diameter, each puncture 2 or 3 times larger than eye facet; pronotal punctures same size as head punctures, separated by a diameter or less, elytral punctures larger than on pronotum, separated by 1-3 times a diameter; metaventral punctures 3 or 4 times as large as on pronotum, separated by a diameter or more medially, becoming larger and contiguous toward lateral margin. Clypeus feebly emarginate apically, lateral angle rounded, clypeus and frons with sparse, long pubescence. Eye canthus elongate, about 8 eye facets long, slightly angled forward, apically acute, reddish yellow. Pronotum narrowed from base to apex, basal angle abruptly rounded, anterior angle evenly rounded, basal margin without bordering line. Epipleuron narrow, grooved, deeply emarginate for reception of femoral apices. Protibia not flanged, outer margin narrowly oblique, smooth. Carinae on prosternal process wide at apex, convergent toward base, acutely joined at base with single carina extended to basal margin of prosternum. Metaventrite without setal tuft. Abdomen with primary pores laterally between ventrites 4- 5 large, deeply indenting apical margin of 4th ventrite; postcoxal line on basal abdominal ventrite rounded, extended to apical margin of ventrite at middle, then broadly forward, ventrite with sparse, long pubescence and dense, sparse punctures medially; ventrites 2-4 sparsely pubescent throughout, punctures small, dense, becoming denser toward lateral margin; 5th ventrite slightly depressed medially, apical margin deeply emarginate, surface densely punctate in basal 1/2, apical 1/2 impunctate; 6th ventrite depressed in apical 1/2, apical margin deeply emarginate, angle on each side of median depression pronounced, abruptly rounded, surface impunctate. Apical tergite finely, indistinctly punctured. Genitalia with basal lobe slightly shorter than paramere, asymmetrical, wide, narrowed and bent in apical 1/6; paramere 
<emphasis id="B952EA83FF8DFF9CFE8259D1FEEBFD60" box="[318,358,644,667]" italics="true" pageId="32" pageNumber="29">Psc</emphasis>
, slender, strongly curved (
<figureCitation id="131D2A14FF8DFF9CFD0C59D6FCB5FD60" box="[688,824,643,667]" captionStart="Figures 86-101. 86-91" captionStartId="80.[192,286,1611,1633]" captionTargetBox="[207,1418,205,1583]" captionTargetId="figure-19@80.[192,1440,192,1618]" captionTargetPageId="80" captionText="Figures 86-101. 86-91) Dilatitibialis gravabilis. 86) Habitus. 87) Phallobase ventral. 88) Phallobase lateral. 89) Enlarged siphonal apex. 90) Sipho. 91) Female genitalia. 92-96) Dilatitibialis lillian. 92) Habitus. 93) Phallobase ventral. 94) Phallobase lateral. 95) Sipho. 96) Enlarged siphonal apex. 97-101) Dilatitibialis tropicalis. 97) Habitus. 98) Phallobase ventral. 99) Phallobase lateral. 100) Sipho. 101) Enlarged siphonal apex." figureDoi="http://doi.org/10.5281/zenodo.5176527" httpUri="https://zenodo.org/record/5176527/files/figure.png" pageId="32" pageNumber="29">Fig. 98, 99</figureCitation>
); sipho robust, strongly curved in basal 2/3, basal capsule large, inner arm short, wide, dorsally truncate, outer arm wide, slightly longer than inner arm, with small accessory piece, basal border deeply emarginate (
<figureCitation id="131D2A14FF8DFF9CFC265996FBB6FD20" box="[922,1083,707,731]" captionStart="Figures 86-101. 86-91" captionStartId="80.[192,286,1611,1633]" captionTargetBox="[207,1418,205,1583]" captionTargetId="figure-19@80.[192,1440,192,1618]" captionTargetPageId="80" captionText="Figures 86-101. 86-91) Dilatitibialis gravabilis. 86) Habitus. 87) Phallobase ventral. 88) Phallobase lateral. 89) Enlarged siphonal apex. 90) Sipho. 91) Female genitalia. 92-96) Dilatitibialis lillian. 92) Habitus. 93) Phallobase ventral. 94) Phallobase lateral. 95) Sipho. 96) Enlarged siphonal apex. 97-101) Dilatitibialis tropicalis. 97) Habitus. 98) Phallobase ventral. 99) Phallobase lateral. 100) Sipho. 101) Enlarged siphonal apex." figureDoi="http://doi.org/10.5281/zenodo.5176527" httpUri="https://zenodo.org/record/5176527/files/figure.png" pageId="32" pageNumber="29">Fig. 100, 101</figureCitation>
).
</paragraph>
<paragraph id="8B993691FF8DFF9CFF4C59B6FE6DFD00" blockId="32.[192,1441,195,763]" box="[240,480,739,763]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF4C59B6FEDEFD00" bold="true" box="[240,339,739,763]" pageId="32" pageNumber="29">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B993691FF8DFF9CFF7C5876FE5BFCC0" blockId="32.[192,470,803,827]" box="[192,470,803,827]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C5876FECFFCC0" bold="true" box="[192,322,803,827]" pageId="32" pageNumber="29">Variation</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33C651AFF8DFF9CFF7C5836FD00FBC0" pageId="32" pageNumber="29" type="materials_examined">
<paragraph id="8B993691FF8DFF9CFF7C5836FE5DFC80" blockId="32.[192,464,867,891]" box="[192,464,867,891]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C5836FEFCFC80" bold="true" box="[192,369,867,891]" pageId="32" pageNumber="29">
<typeStatus id="549D8833FF8DFF9CFF7C5836FE8EFC80" box="[192,259,867,891]" pageId="32" pageNumber="29">Type</typeStatus>
locality
</emphasis>
. 
<collectingCountry id="F3317601FF8DFF9CFEC35836FE41FC80" box="[383,460,867,891]" name="Brazil" pageId="32" pageNumber="29">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8B993691FF8DFF9CFF7C58F6FC8EFC40" blockId="32.[192,771,931,955]" box="[192,771,931,955]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C58F6FE16FC40" bold="true" box="[192,411,931,955]" pageId="32" pageNumber="29">Type depository</emphasis>
. 
<collectionCode id="ED37AE54FF8DFF9CFE1458F6FE74FC40" box="[424,505,931,955]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="32" pageNumber="29">ZMUC</collectionCode>
(
<typeStatus id="549D8833FF8DFF9CFDBB58F6FDFEFC40" box="[519,627,931,955]" pageId="32" pageNumber="29" type="holotype">holotype</typeStatus>
, examined).
</paragraph>
<paragraph id="8B993691FF8DFF9CFF7C58B6FC35FC00" blockId="32.[192,952,995,1019]" box="[192,952,995,1019]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C58B6FDAAFC00" bold="true" box="[192,551,995,1019]" pageId="32" pageNumber="29">Geographical distribution</emphasis>
. Known only from the Brazilian.
</paragraph>
<paragraph id="8B993691FF8DFF9CFF7C5F76FD00FBC0" blockId="32.[192,653,1059,1083]" box="[192,653,1059,1083]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C5F76FE51FBC0" bold="true" box="[192,476,1059,1083]" pageId="32" pageNumber="29">Specimens examined</emphasis>
. The 
<typeStatus id="549D8833FF8DFF9CFDA25F76FD04FBC0" box="[542,649,1059,1083]" pageId="32" pageNumber="29" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C651AFF8DFF9CFF7C5F36FBECFB20" pageId="32" pageNumber="29" type="discussion">
<paragraph id="8B993691FF8DFF9CFF7C5F36FE6AFB60" blockId="32.[192,1440,1123,1243]" pageId="32" pageNumber="29">
<emphasis id="B952EA83FF8DFF9CFF7C5F36FEB7FB80" bold="true" box="[192,314,1123,1147]" pageId="32" pageNumber="29">Remarks</emphasis>
. The dorsal color pattern is similar to that of several other species of 
<emphasis id="B952EA83FF8DFF9CFB355F36FAA0FB80" box="[1161,1325,1123,1147]" italics="true" pageId="32" pageNumber="29">
<taxonomicName id="4C264D12FF8DFF9CFB355F36FAA5FB80" authorityName="Duverger" authorityYear="2001" box="[1161,1320,1123,1147]" class="Insecta" family="Coccinellidae" genus="Dilatitibialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="29" phylum="Arthropoda" rank="genus">Dilatitibialis</taxonomicName>
,
</emphasis>
but male genitalia are distinctive.
</paragraph>
<paragraph id="8B993691FF8DFF9CFF4C5FF6FBECFB20" blockId="32.[192,1440,1123,1243]" pageId="32" pageNumber="29">
<bibRefCitation id="EFB74B60FF8DFF9CFF4C5FF6FE26FB40" author="Mulsant, M. E." box="[240,427,1187,1211]" pageId="32" pageNumber="29" pagination="1 - 1104" refId="ref57076" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant (1850)</bibRefCitation>
had only one example of this species, the 
<typeStatus id="549D8833FF8DFF9CFC0C5FF6FB95FB40" box="[944,1048,1187,1211]" pageId="32" pageNumber="29" type="holotype">holotype</typeStatus>
male labeled “ 
<collectingCountry id="F3317601FF8DFF9CFB715FF6FA96FB40" box="[1229,1307,1187,1211]" name="Brazil" pageId="32" pageNumber="29">Brasil</collectingCountry>
. Tropicalis Muls. (handwritten)/ 
<typeStatus id="549D8833FF8DFF9CFE0A5F96FDD5FB20" box="[438,600,1219,1243]" pageId="32" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C264D12FF8DFF9CFDDC5F96FC31FB20" authority="Mulsant" authorityName="Mulsant, 1850:" authorityYear="1038" box="[608,956,1219,1243]" class="Insecta" family="Coccinellidae" genus="Cleothera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="29" phylum="Arthropoda" rank="species" species="tropicalis">Cleothera tropicalis Mulsant</taxonomicName>
(red paper).”
</paragraph>
</subSubSection>
</treatment>
</document>