<document id="7E561CC55905B7665F3C06F653B48A88" ID-DOI="10.5252/g2011n2a10" ID-ISSN="1638-9395" ID-Zenodo-Dep="5372482" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630272317791" checkinUser="marcus" docAuthor="Delfino, Massimo, Bailon, Salvador &amp; Pitruzzella, Gaetano" docDate="2011" docId="038387F3FFCCDE03382CFD78B57CFBCE" docLanguage="en" docName="Geodiversitas.2011.33.2.357-382.pdf" docOrigin="Geodiversitas 33 (2)" docSource="http://dx.doi.org/10.5252/g2011n2a10" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Amphisbaenia Gray 1844" docType="treatment" docVersion="3" lastPageNumber="373" masterDocId="FFBAFF8BFFDCDE133920FF8AB10BFFE5" masterDocTitle="The Late Pliocene amphibians and reptiles from “ Capo Mannu D 1 Local Fauna ” (Mandriola, Sardinia, Italy)" masterLastPageNumber="382" masterPageNumber="357" pageNumber="373" updateTime="1699207784534" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="FD47E3F311A8D3D752FF748CDAED9786">The Late Pliocene amphibians and reptiles from “ Capo Mannu D 1 Local Fauna ” (Mandriola, Sardinia, Italy)</mods:title>
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<mods:namePart id="25A38B529B312ECBE1A0FF536DF08DFF">Delfino, Massimo</mods:namePart>
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<mods:namePart id="5232E152988B4BB64D4289447D15172B">Bailon, Salvador</mods:namePart>
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<mods:namePart id="7F4B1A2FE00AD9DFB674A58E7107085A">Pitruzzella, Gaetano</mods:namePart>
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<taxonomicName id="4C2A4D66FFCCDE03382CFD78B0ADFCE8" authorityName="Gray" authorityYear="1844" box="[268,422,754,781]" order="Amphisbaenia" pageId="16" pageNumber="373" rank="order">Amphisbaenia</taxonomicName>
indet. (
<figureCitation id="13112A60FFCCDE033865FC98B088FCC8" box="[325,387,786,813]" captionStart="FIG" captionStartId="18.[113,124,1380,1397]" captionTargetBox="[150,1173,221,1337]" captionTargetId="figure-184@18.[614,1082,577,942]" captionTargetPageId="18" captionText="FIG. 6. — Fossil remains of reptiles from Capo Mannu D1 Local Fauna (Sardinia): A-C, Amphisbaenia indet., trunk vertebra (DSTC 6021), dorsal, left lateral and ventral views; D-F, Eryx cf. E. jaculus, trunk vertebra (DST 6022), dorsal, ventral and posterior views; G, caudal vertebra (DSTC 6023), anterior view. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5372496" httpUri="https://zenodo.org/record/5372496/files/figure.png" pageId="16" pageNumber="373">Fig. 6</figureCitation>
A-C)
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<subSubSection id="C330656EFFCCDE033951FCC6B1EAFC9B" pageId="16" pageNumber="373" type="materials_examined">
<paragraph id="8B9536E5FFCCDE033951FCC6B1EAFC9B" blockId="16.[113,646,843,894]" pageId="16" pageNumber="373">
<collectionCode id="ED3BAE20FFCCDE033951FCC6B1DDFC87" box="[113,214,844,867]" pageId="16" pageNumber="373">MATERIAL</collectionCode>
EXAMINED. — Trunk vertebra: 16; caudal vertebra: 1.
</paragraph>
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<subSubSection id="C330656EFFCCDE033951FC3BB377F9C0" pageId="16" pageNumber="373" type="description">
<paragraph id="8B9536E5FFCCDE033951FC3BB009FC2C" blockId="16.[111,648,945,1573]" box="[113,258,945,971]" pageId="16" pageNumber="373">DESCRIPTION</paragraph>
<paragraph id="8B9536E5FFCCDE033951FC5AB0CDFA42" blockId="16.[111,648,945,1573]" pageId="16" pageNumber="373">
The trunk vertebrae have a maximum centrum length of 
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. They show a rather uniform morphology with distinctly dorsoventrally depressed cotyles and condyles; wide oval-shaped prezygapophyses characteristically “protruding” from the rest of the vertebra in dorsal view and bearing large and short blunt processes; well-defined globular synapophyses; convex anterior and concave posterior edge of the neural arch; underdeveloped neural crest (in some cases a hint of a ridge terminating in a flattened area can be present close to the posterior end of the neural arch); depressed neural arch (when seen in posterior view); flattened ventral surface of the centrum with parallel lateral margins and two small foramina in the anterior region.
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<paragraph id="8B9536E5FFCCDE0339A8FA21B377F9C0" blockId="16.[111,648,945,1573]" pageId="16" pageNumber="373">The caudal vertebra has a similar morphology but differs for the absence of prezygapophyseal processes and for the presence of anterior transverse and posterior haemal processes fused to the centrum.</paragraph>
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<subSubSection id="C330656EFFCCDE033951F9C0B57CFBCE" pageId="16" pageNumber="373" type="discussion">
<paragraph id="8B9536E5FFCCDE033951F9C0B1F8F987" blockId="16.[113,647,1610,1700]" box="[113,243,1610,1636]" pageId="16" pageNumber="373">DISCUSSION</paragraph>
<paragraph id="8B9536E5FFCCDE033951F9E3B38CF941" blockId="16.[113,647,1610,1700]" pageId="16" pageNumber="373">The combination of the morphological characters described above allows to identify a member of the</paragraph>
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<taxonomicName id="4C2A4D66FFCCDE033B83FF52B235FF16" authorityName="Gray" authorityYear="1844" box="[675,830,216,243]" order="Amphisbaenia" pageId="16" pageNumber="373" rank="order">Amphisbaenia</taxonomicName>
, but due to the fact that isolated amphisbaenian vertebrae are generally not diagnostic even at genus level, a precise taxonomic allocation is not possible (see 
<bibRefCitation id="EFBB4B14FFCCDE033A5EFEBDB51CFEB7" author="DELFINO M." box="[894,1047,311,338]" pageId="16" pageNumber="373" pagination="89 - 97" refId="ref14766" refString="DELFINO M. 1997. - Blanus from the Early Pleistocene of Southern Italy: another small tessera from a big mosaic, in BOHME W., BISCHOFF W. &amp; ZIEGLER T. (eds), Herpetologia Bonnensis. Societas Herpetologica Europaea, Bonn: 89 - 97." type="book chapter" year="1997">Delfino 1997</bibRefCitation>
, and literature therein). It is however likely that these vertebrae belong to 
<taxonomicName id="4C2A4D66FFCCDE033A31FEFCB51BFE74" authority="Kearney, 2003" authorityName="Kearney" authorityYear="2003" box="[785,1040,374,401]" class="Reptilia" family="Blanidae" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="373" phylum="Chordata" rank="family">Blanidae Kearney, 2003</taxonomicName>
, the only family that has been identified so far in the European fossil record. The fossil remains are morphologically congruent with the extant genus 
<taxonomicName id="4C2A4D66FFCCDE033AFEFE5FB5B1FE15" authority="Wagler, 1830" authorityName="Wagler" authorityYear="1830" box="[990,1210,469,496]" class="Reptilia" family="Blanidae" genus="Blanus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="B95EEAF7FFCCDE033AFEFE5FB52FFE0A" box="[990,1060,469,495]" italics="true" pageId="16" pageNumber="373">Blanus</emphasis>
Wagler, 1830
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and not with, for example, 
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<emphasis id="B95EEAF7FFCCDE033AE6FE7FB5B2FDEA" box="[966,1209,501,527]" italics="true" pageId="16" pageNumber="373">Trogonophis wiegmanni</emphasis>
Kaup, 1830 (
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whose vertebrae are comparatively more elongated and narrow.
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Noteworthy is that 
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listed the presence of 
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<emphasis id="B95EEAF7FFCCDE033A67FDFEB2D2FD6B" box="[839,985,628,654]" italics="true" pageId="16" pageNumber="373">Blanus gracilis</emphasis>
(Roček, 1984)
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in the Lower Miocene Sardinian site of Oschiri. This species was originally described on the basis of isolated vertebrae which are notoriously poorly informative (
<bibRefCitation id="EFBB4B14FFCCDE033B8BFD78B214FCE8" author="RAGE J. - C." box="[683,799,754,781]" pageId="16" pageNumber="373" pagination="3 - 27" refId="ref16001" refString="RAGE J. - C. 1988. - Le gisement du Bretou (Phosphorites du Quercy, Tarn-et-Garonne, France) et sa faune de vertebres de l'Eocene superieur. I. Amphibiens et Reptiles. Palaeontographica (A) 205: 3 - 27." type="journal article" year="1988">Rage 1988</bibRefCitation>
; 
<bibRefCitation id="EFBB4B14FFCCDE033A08FD78B2BCFCE8" author="DELFINO M." box="[808,951,754,781]" pageId="16" pageNumber="373" pagination="89 - 97" refId="ref14766" refString="DELFINO M. 1997. - Blanus from the Early Pleistocene of Southern Italy: another small tessera from a big mosaic, in BOHME W., BISCHOFF W. &amp; ZIEGLER T. (eds), Herpetologia Bonnensis. Societas Herpetologica Europaea, Bonn: 89 - 97." type="book chapter" year="1997">Delfino 1997</bibRefCitation>
; 
<bibRefCitation id="EFBB4B14FFCCDE033AE0FD78B538FCE8" author="AUGE M." box="[960,1075,754,781]" pageId="16" pageNumber="373" pagination="1 - 369" refId="ref14246" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Histoire naturelle 192: 1 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
) but it seems that tooth bearing bones are present at Oschiri and that therefore a valid diagnosis of this taxon based on diagnostic characters could be possible. In any case, the absence of tooth bearing elements in the Capo Mannu D1 LF does not allow confirmation of the possible survival of this taxon in 
<collectingRegion id="49EEF807FFCCDE033D20FC3AB550FC2E" box="[1024,1115,944,971]" country="Italy" name="Sardegna" pageId="16" pageNumber="373">Sardinia</collectingRegion>
.
</paragraph>
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Order 
<taxonomicName id="4C2A4D66FFCCDE033A0FFB9AB57CFBCE" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[815,1143,1040,1067]" order="Serpentes" pageId="16" pageNumber="373" rank="order">SERPENTES Linnaeus, 1758</taxonomicName>
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