Tryphosa nana ( Krøyer, 1846a )

Kilgallen, N. M. & Lowry, J. K., 2014, The Tryphosa group (Crustacea: Amphipoda: Lysianassoidea: Lysianassidae: Tryphosinae), Zootaxa 3768 (5), pp. 501-545 : 537-539

Tryphosa nana ( Krøyer, 1846a)

( Fig. 23 )

Anonyx nanus Krøyer, 1846a: 30 .― Krøyer, 1846b: pl. 17 fig. 2a–t.― Bruzelius, 1859: 42.― Bate, 1862: 78, pl. 12 fig. 2. ― Lilljeborg, 1865a: 18.― Lilljeborg, 1865b: 28.― Heller, 1866: 24.― Norman, 1869: 274.― Stalio, 1877: 243.― Graeffe, 1880: 48.― Stossich, 1880: 71.― Nebeski, 1881: 34.― Krapp-Schickel, 1974: 321, 338.

Tryphosa nanus .― Boeck, 1871: 118.― Boeck, 1876: 181.

Tryphosa ciliata Sars, 1883: 81 , pl. 3 fig. 4.― Chevreux, 1887: 295, 331.― Walker, 1888: 172, pl. 13 figs 1–4.― Walker, 1889: 77.― Walker, 1892: 241.

Tryphosa nana .― Chevreux, 1888: 3.― Meinert, 1890: 155.― Bonnier, 1893: 191, 196, pl.7.― Norman, 1900: 203.― Scott, 1901.― Christodoulou, Paraskevopoulou, Syranidou & Koukouras, 2013: 1312, table 2.

Tryphosa höringii .― Walker, 1889: 77.

Orchomenella ciliata .― Sars, 1891: 69, pl. 25 fig. 2.― Robertson, 1892: 204.― Sars, 1895: 683.― Walker, 1895a: 420. ― Walker, 1895b: 292.

Anonyx pinguis .― Della Valle, 1893: 821, pl. 28 figs 22–35.

Orchomenella nana .― Chevreux, 1898: 476.― Walker, 1898: 166.― Walker, 1904: 244.― Stephensen, 1923: 12.―Chevreux, 1925: 288.― Chevreux & Fage, 1925: 71, fig. 62.― Stephensen, 1929: 60.―Marine Biological Association of the UK, 1931: 188.― Stephensen, 1932: 357.― Stephensen, 1935: 104 (key).― Crawford, 1937: 644.― Moore, 1937: 117. ― Schellenberg, 1942: 108.― Stephensen, 1942: 474.― Gorbunov, 1946: 43.― Jones, 1948: 404.― Enequist, 1949: 387, 400.― Reid, 1951: 195.―J.L. Barnard, 1958: 97.― Oldevig, 1959: 22.― Duhig, 1960: 64.― Agrawal, 1963: 632, pl. 3. ― Gurjanova, 1964: 279.―Toulmond, 1964: 321.― Toulmond & Truchot, 1964: 7.― Agrawal, 1966: 459, pl. 5.― Nayar, 1966: 135.― O'Riordan, 1969: 32.― Bellan-Santini & Ledoyer, 1973: 920.― Laverack & Blackler, 1974: 81.― Drago & Albertelli, 1978: 212.― Myers & Southgate, 1980: 975.― Walker & Rees, 1980: 56.― Macquart-Moulin, 1984: 185. ― Diviacco & Ruffo, 1989: 525, fig. 359 (in part).— Dauvin & Sorbe, 1995: 443, 447, 449, 456, tables 4, 5.— Stefanidou & Voultsiadou-Koukoura, 1995: 602, table 1.— Cartes & Sorbe, 1999: 1138, table 1, 1134, table 4.— Bakir & Çevirgen, 2010: 83, table 1.

Orchomenella nanus .― Stebbing, 1906: 81.― Tattersall, 1913: 4.― Schellenberg, 1925: 118.

Orchomene nana .―J.L. Barnard, 1964: 89 (key).― Karaman, 1973: 133, figs 14–18 (in part).― Krapp-Schickel, 1974: 321, 338.― Kitching et al, 1976: 756.― Lincoln, 1979: 70, figs 25a–g, 26e–h.― O'Riordan, 1982: 551.― Moore, 1984: 35. ― Parker, 1984: 122.― Bellan-Santini, 1985: 275.― Marques & Bellan-Santini, 1985: 323, 349.― Holmes, 1986: 37. ― Costello et al, 1989: 34.― Kaartvedt, 1989: 189, table 3, 191, table 5.― Dauvin & Gentil, 1990: 126, 128.― Buhl-Jensen & Fosså, 1991: 248.― Hamerlynck & Mees, 1991: 208. table 2.― Mees & Hamerlynck, 1992: 360, table 2.― Cattrijsse, Mees & Hamerlynck, 1993: 191, table 1.― Marques & Bellan-Santini, 1993: 557, table 2, 559, table 3, 560, table 4, 561, table 5.― Dauvin et al, 1994: 550, table 3.― Dauvin & Zouhiri, 1996: 914, table 3, 916, table 5, 921, table 6.― Conradi & Lopez-Gónzález, 1999: 6, table 1.

Orchomene nanus .― Barnard & Karaman, 1991: 509.― Conlan, 1994: 520

not Tryphosa nana .― Sars, 1891: 76, pl. 27 fig. 1 (= Tryphosella sarsi ).― Sars, 1895: 684 (= Tryphosella sarsi ).―Walker, 1895a: 422 (= Tryphosella sarsi ).― Walker, 1895b: 294 (= Tryphosella sarsi ).

not Tryphosa nanus .― Robertson, 1892: 205 (?= Tryphosella sarsi ).

not Orchomene nana .― Karaman, 1973: 133, figs 14–18 (in part = Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).

not Orchomenella nana .― Ledoyer, 1977: 384 (= Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).― Diviacco & Ruffo, 1989: 525, fig. 359 (in part = Orchomenella crenata Chevreux & Fage, 1925 according to Kilgallen et al. 2006).

Types. According to Dr Jørgen Olesen ( ZMUC) there are two sets of material ( ZMUC CRU-7476 ?type and ZMUC CRU-7487 syntype) which might be considered as original material .

Material examined. 23 specimens & 1 slide (female, 5.5 mm), SMNH N. Amph. 6904, from Gåsörännan, Sweden, 30–40 m ; 8 specimens, AM P.11222, from Plymouth, Devon, United Kingdom (50°24'N 4°7'W), coll. A.A. Cameron.

Type locality. The Kattegat.

Habitat. Sand, mud and shelly debris.

Feeding strategies. Often found ‘stripping bait in lobster pots’ ( O’Riordan 1982) on carcasses of dead crabs ( Norman 1900, Moore 1984), also in Echinocardium tests ( Laverack & Blackler 1974).

Depth range. 0–250 m ( Lincoln 1979).

Distribution. Tryphosa nana is widely recorded and locally very common along the Atlantic coasts of Europe and the North Sea, from Sweden and southern Norway to Spain ( Chevreux & Fage 1925); also reported from Senegal ( Schellenberg 1925) and Guinea ( Reid 1951). It is also recorded from the Adriatic Sea ( Heller 1866, Karaman 1973), Aegean Sea ( Bellan-Santini 1985, Christodoulou et al. 2013) and Mediterranean Sea (Bellan- Santini & Ledoyer 1973).

Figures

FIGURE 23. Tryphosa nana (Krøyer, 1846), female, 5.6 mm, SMNH N. Amph. 6904, Gåsörännan, Sweden; female whole animal and head of male, after Sars, 1891 (as Orchomenella ciliata). Scale bars: gnathopods, pereopods, 0.2 mm; remainder, 0.1 mm.