Toxognathus costulatus Fairmaire, 1878

Rosa, Simone Policena, 2014, Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae), Papéis Avulsos de Zoologia (São Paulo) 54 (18), pp. 217-292 : 242

Toxognathus costulatus Fairmaire, 1878

( Figs. 9 , 10 , 34I )

Toxognathus costulatus Fairmaire, 1878: 271 ; Schenkling, 1927: 509; Fleutiaux, 1918a: 276; Fleutiaux, 1940a: 35.

Redescription (male, Fig. 34I ): Integument dark brown or dark reddish-brown. Total length 11-12 mm; elytral base as wide as pronotum, elytra 2.7-3.0x longer than pronotum. Frons ( Figs. 9D, E ) concave medially, frontal carina absent medially. Antennae ( Fig. 9A ) surpassing posterior angles of pronotum by 2 antennomeres. Mandible unidentate ( Fig. 9D ). Pronotum ( Fig. 9F ) flattened 1.15-1.30x wider than long with sides sinuous converging abruptly anteriad on anterior half, lateral carina directed ventrad anteriorly, posterior angles divergent and convex; with punctures less than 0.5 diameter apart. Elytral interstices costate. Free margin of metacoxal plate short and triangular; tibiae widened apicad, dorso apical angle of protibiae acute and produced ( Fig. 9J ). Tergite IX ( Fig. 10D ) with anterior margin bisinuous.

Aedeagus ( Figs. 10E, F ): Phallobase 0.44x total length of aedeagus, 1.3x longer than wide, length ratio between lateroposterior and median parts 3.1; parameres fused to each other on ventral anteromedian margin; basal strut 0.34x total length of penis; ventral sclerite elliptical.

Holotype: [Conchinch. (illegible) 1877] [ Toxognathus gen. et sp. nov. costulatus, Fairm., Conchonchine ] [ Toxognathus costulatus Fairm., J. Chassain det. 05] [Muséum Paris, Coll. E. Fleutiaux] male, (MNHN).

Material examined: VIETNAM. 1 ex. ( MNHN). Tay Ninh , 2 exs. ( MNHN) .

Distribution: VIETNAM. Tay Ninh.

Remarks: Toxognathus costulatus is easily recognized by its protibia widened apicad with dorso apical angle acute and produced ( Fig. 9J ). The parameres with anteromedian margin fused together ventrally is unique to this species.

Figures

FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate.

FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral).

FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes.