Tipula (Vestiplex) coronifera Savchenko

Starkevich, Pavel, Podenas, Sigitas & Gelhaus, Jon K., 2020, Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia, Zootaxa 4837 (1), pp. 1-88 : 32-35

publication ID	https://doi.org/ 10.11646/zootaxa.4837.1.1
publication LSID	lsid:zoobank.org:pub:001B5DF8-A24D-4353-8BEA-8B181BE9D279
DOI	https://doi.org/10.5281/zenodo.4454917
persistent identifier	https://treatment.plazi.org/id/03E187A2-9339-A848-6FEB-FB9A21CDC155
treatment provided by	Plazi
scientific name	Tipula (Vestiplex) coronifera Savchenko
status

Treatment

( Figs 25 View FIGURES 14–27 , 50–62 View FIGURES 50–57 View FIGURES 58–62 ; Map 3 View MAPS 1–3 )

Tipula (Vestiplex) coronifera Savchenko, 1960: 210 ; 1964: 201; Oosterbroek & Theowald, 1992: 154.

Type material examined. HOLOTYPE ♂: MONGOLIA, Khangai Mts. (south-east), Lamyn-Gegen, 16 VII 1926 (Kirichenko), Nr 750 ( ZIN) . PARATYPE: ♀, Nr 770, topotypic ( ZIN) .

Additional material examined. MONGOLIA. Uvs Aimag: 1 ♀, Khovd Soum, Shiver / Ireg Gol , 25 km NE Khovd, N49.43119, E91.19093, 1964 m, 5–6.vii.2010, MAIST, MAIS2010070504 GoogleMaps . Hovsgol Aimag: 1 ♂, Ulaan-Uul Soum, unnamed trib. of Beltes Gol 34 km NE Bayanzurh, 2113 m, N50.42980, E99.21690, 28.vi.2006, J. Gelhaus #1054-A, SRP06062803 GoogleMaps . Ovorhangay Aimag: 1 ♀, Zuunbayan-Ulaan Soum , Millenium Road A 0301 marker 380.6 km ~ 50 km NE Arvayheer, N46.60919, E103.10400, 1820 m, 5.vii.2004, SRPT, SRP04070502 GoogleMaps .

Elevation range in Mongolia. Adults were collected at altitudes ranging from 1960 m to 2100 m.

Period of activity. Adults are active from the end of June through to early July.

Known distribution. Mongolia ( Map 3 View MAPS 1–3 ).

Redescription. Male. Body length 20.4 mm, wing length 20.8 mm. General body color brownish yellow.

Head. Brownish yellow, vertex and occiput brown gray pruinose with brown median line. Rostrum brownish yellow, nasus distinct. Antenna 13-segmented, if bent backward reaching base of wing. Scape and pedicel yellow, first flagellomere yellow, darkened brown mesally. Flagellomeres 2–6 brownish, succeeding flagellomeres yellowish, apical flagellomere brownish, small, reduced, distinctly shorter than preceding flagellomere. Each flagellomere, except first, with darkened basal enlargement and moderately incised. Verticils shorter than respective segments. Palpus brown.

Thorax. Brownish gray. Pronotum brownish yellow with brown median line. Prescutum and presutural scutum brown with 4 darker longitudinal stripes bordered by brown, intermediate stripes completely separated from each other. Interspace between median and lateral stripes yellowish with pale setae arising from dark sockets. Postsutural scutum brown. Scutal lobe with 2 indistinct spots bordered by brown. Scutellum brownish, postnotum brownishyellow, with brown median line. Pleura brown, sparsely dusted with gray. Coxae yellow, sparsely dusted with grey. Trochanters, femora and tibiae yellow. Tarsal segments indistinctly passing into brownish. Distal part of femora and tibiae narrowly darkened. Tarsal claws without tooth. Wing indistinctly patterned. Halter with pale stem and brown knob.

Abdomen. Yellow; tergite 1 laterally darkened and sparsely pruinose. Dorsal median stripe indistinct, interrupted at caudal margins of segments. Lateral stripe brownish and distinct. Lateral margins of tergites pale. Sternite 1 sparsely dusted with gray.

Hypopygium. Brown, at base slightly broader than abdomen. Tergite 9 in shape of large concave crescent sclerotised saucer ( Fig. 50 View FIGURES 50–57 ). Main body of tergal saucer brownish yellow with blackened rim. Posterior margin of tergal saucer broadly emarginated, with small denticles or teeth. Median V-shaped notch inconspicuous. Lateral angle of tergal saucer with broad obtuse tooth. Anterior and lateral portions of tergal saucer raised into sclerotised border. Border anteriorly inconspicuously toothed, laterally produced into acute tooth directed caudad and situated under lateral angle of tergal saucer so tergite 9 with 2 teeth in lateral view. Gonocoxite unarmed ( Fig. 51 View FIGURES 50–57 ). Outer gonostylus flattened, nearly oval ( Fig. 52 View FIGURES 50–57 ). Inner gonostylus in shape of claw-shaped plate, with 2 mid-dorsal teeth, anterior tooth twice as large as posterior. Beak extended into blackened, obtuse rostrum ( Fig. 53 View FIGURES 50–57 ). Gonocoxal fragment large with lateral and medial sclerites well-developed ( Fig. 54 View FIGURES 50–57 ). Medial sclerites fused, anterior apodeme long and broad, posterior part flat and arched. Lateral sclerite large and bilobed. Aedeagal guide in shape of long, slender tube ( Fig. 55 View FIGURES 50–57 ). Sperm pump with central vesicle small and flattened ( Fig. 56 View FIGURES 50–57 ). Compressor apodeme with median incision. Posterior immovable apodeme much longer than compressor apodeme, large and extended laterally into broad plate. Anterior immovable apodeme in shape of rounded plate. Aedeagus shaped as very long tube similar to T. (V.) subcentralis . Distal part ventrally membranous, shovel-shaped ( Fig. 57 View FIGURES 50–57 ).

Female. Body length 26.8–27.0 mm, wing length 22.0– 26.6 mm. Antenna short, flagellomeres except first with very small incisions. Mesopleura brown, gray pruinose. Distal part of femora darkened.

Ovipositor ( Figs 58–62 View FIGURES 58–62 ). Tergite 10 shiny, brown. Cercus brownish yellow, slender, about as long as tergite 10, with tip narrowed, outer margin almost smooth with small serration ( Fig. 58 View FIGURES 58–62 ). Hypovalva in shape of pale, elongated slender filament, with short trichia at tip ( Fig. 59 View FIGURES 58–62 ). Median incision between hypovalvae deeper than posterior margin of sternite 8. Lateral incision relatively deep and narrow (about twice as deep as maximal width). Lateral angle of sternite 8 rounded. Sternite 9 with anterior part slightly broadened, posterior part rounded ( Fig. 60 View FIGURES 58–62 ). Furca anteriorly narrowed, posteriorly broad ( Fig. 61 View FIGURES 58–62 ). Bursa copulatrix with spermathecal ducts sclerotised at base, in shape of lightly curved, slender dark brown process. Wall of bursa copulatrix on connection site with spermathecal ducts partially sclerotised, sclerotisation extended short distance along wall ( Fig. 62 View FIGURES 58–62 ). Cul-de-sac of bursa copulatrix curved. Spermatheca spherical, greatly broadened at base ( Fig. 25 View FIGURES 14–27 ).