Thoracostrongylus formosanus flavipes, Xia & Tang & Schillhammer, 2022

Xia, Mei-Hua, Tang, Liang & Schillhammer, Harald, 2022, Review of Chinese species of the genus Thoracostrongylus Bernhauer, 1915 (Coleoptera, Staphylinidae, Staphylininae), ZooKeys 1131, pp. 99-134 : 99

Thoracostrongylus formosanus flavipes ssp.nov.

Figs 46-52 , 113 台湾钝胸隐翅虫黄足亚种

Type material.

Holotype. China - Zhejiang Prov. • ♂, glued on a card with labels as follows: "China: Zhejiang, Longquan, Fengyang Mt., Guanyintai; alt. 1000 m; 11 May 2019; Tang & Zhao leg." "Holotype / Thoracostrongylus formosanus flavipes / Xia, Tang & Schillhammer" [red handwritten label]; SHNU. Paratypes. China - Zhejiang Prov. • 4♂♂, 6♀♀; same data as holotype; SHNU • 1♂; Longquan City, Fengyangshan, Lu’ao Vill.; 27°55'8.95"N, 119°11'55.54"E; alt. 1200-1300 m; 16-17 July 2018; Zi-Wei Yin leg.; SHNU • 2♂♂, 1♀; Longquan City, Fengyangshan N.R., Lu’aocun Village; 27°55'19.66"N, 119°11'38.86"E; alt. 1076 m; 04 May 2016; Jiang, Liu & Zhou leg.; SHNU • 1♂; Longquan City, Fengyangshan N.R., Datianping; 27°54'29.67"N, 119°10'31.45"E; alt. 1350 m; 30 April 2016; Jiang, Liu & Zhou leg.; SHNU • 2♀♀; Longquan City, Fengyangshan N.R., Datianping; 27°54'29.67"N, 119°10'31.45"E; alt. 1350 m; 30 April 2016; Jiang, Liu & Zhou leg.; SHNU • 2♂♂, 1♀; Jinhua City, Pan’an County, Dapanshan N.R.; 28°58'41.03"N, 120°29'19.24"E; alt. 531-783 m; 08 May 2016; Jiang, Liu & Zhou leg.; SHNU • 1♂, 2♀♀; Lishui City, Qingyuan County, Baishanzu, Station to Peak; 27°45'20"N, 119°11'78"E; alt. 1721 m; 24 April 2015; Song & Yan leg.; SHNU • 2♀♀; Longquan City, Fengyangshan N.R., Mihougu, near stream; 27°55'0.18"N, 119°11'52.91"E; alt. 1116 m; 03 May 2016; Jiang, Liu & Zhou leg.; SHNU • 1♀; Wuyanling; alt. 700 m; 09 May 2004; Hu, Tang & Zhou leg.; SHNU. - Guangxi Prov. • 7♂♂, 16♀♀; Huanjiang, Jiuwan Mt., Yangmeiao; 25°12'22.15"N, 108°40'32.01"E; alt. 1250 m; 25 April 2021; Tang, Peng, Cai & Song leg.; SHNU • 1♂; Huanjiang, Jiuwan Mt., Yangmeiao; 25°12'22.15"N, 108°40'32.01"E; alt. 1250 m; 08 May 2021; Tang, Peng, Cai & Song leg.; SHNU • 1♀; Huanjiang, Jiuwan Mt., Yangmeiao; 25°12'22.15"N, 108°40'32.01"E; alt. 1250 m; 23 April 2021; Tang, Peng, Cai & Song leg.; SHNU • 1♂; Jinxiu County, Mt. Shengtangshan; alt. 1500 m; 26 July 2011; Zhong Peng leg.; SHNU • 1♀; Guilin City, Huaping N.R., Anjiangping; alt. 1500 m; 18 July 2011; Liang Tang leg.; SHNU. - Guangdong Prov. • 4♂♂, 2♀♀; Ruyuan County, Nanling N.R., Qingshui Valley; 24°54'57"N, 113°01'55"E; alt. 900 m; 04 May 2015; Peng, Tu & Zhou leg.; SHNU • 1♂, 2♀♀; Ruyuan County, Nanling N.R., Laopengkeng; 24°56'29"N, 113°00'27"E; alt. 1360 m; 29 April 2015; Peng, Tu & Zhou leg.; SHNU • 6♂♂, 3♀♀; Ruyuan County, Nanling N.R., Baobaoshan Station; 24°55'43"N, 113°00'58"E; alt. 1030 m; 25 April 2015; Peng, Tu & Zhou leg.; SHNU. - Sichuan Prov. • 3♂♂, 1♀; Dayi County, Xiling Snow Mt.; 30°38'6.25"N, 103°10'99.08"E; alt. 1250 m; 31 July 2021; Zhao & Cai leg.; SHNU • 1♀; Jiulong County, Hongba; alt. 2000 m; 13 August 2005; Ming Yi leg.; SHNU. - Anhui Prov. • 1♀; Huangshan, Tangkou Town, Hougu; 30°05'3.48"N, 118°08'45.96"E; alt. 569-688 m; 29 June-03 July 2020; Chong Li leg.; SHNU. - Jiangxi Prov. • 1♂, 1♀; Yichun City, Fengxin County, Baizhang Vill.; 28°42'55"N, 114°46'14"E; alt. 1000-1300 m; 16 July 2013; Hu & Lv leg.; SHNU • 1♂; Longnan County, Jiulianshan, summit of Huangniushi; 24°30'53"N, 114°26'6.72"E; alt. 1000-1230 m; 12 May 2021; Zhou & Li leg.; SHNU • 1♂; Ji’an City, Jinggangshan, Huangyangjie; 26°37'25"N, 114°06'58"E; alt. 1240 m; 28.vii,2014; Chen, Hu, Lv & Yu leg.; SHNU • 1♀; Pingxiang City, Gaozhou County, Gaotianyan; 27°23'51"N, 114°00'54"E; alt. 1025 m; 23 July 2013; Song, Yin & Yu leg.; SHNU. - Hunan Prov. • 2♂♂; Liuyang City, Daweishan; 28°25'25"N, 114°05'57"E; alt. 1300 m; 06 June 2014; Peng, Shen, Yu & Yan leg.; SHNU • 2♂♂; Liuyang City, Daweishan, 28°25'25"N, 114°05'57"E; alt. 1300 m, 07 June 2014; Peng, Shen, Yu & Yan leg.; SHNU • 2♂♂, 1♀; Yanling County, Nanfengmian; 26°18'N, 114°01'E; alt. 1855 m; 07 June 2015; Peng, Shen, Tu & Zhou leg.; SHNU • 1♂; Xin’ning County, Shunhuang Mt., Yangheping; 26°23'41.58"N, 111°00'08.16"E; alt. 820 m; 02 May 2021; Yin, Zhang, Pan & Shen leg.; SHNU • 1♂; Xin’ning County, Shunhuang Mt., Yangheping; 26°23'41.58"N, 111°00'08.16"E; alt. 820 m; 30 April 2021; Yin, Zhang, Pan & Shen leg.; SHNU • 1♂; Chengzhou, Yizhang County, Mangshan N.R.; 24°56'26"N, 112°59'18"E; alt. 1400 m; 26 April 2015; Peng, Tu & Zhou leg.; SHNU • 2♀♀; Liuyang City, Daweishan; 28°25'N, 114°05'E; alt. 1000 m; 11 June 2015; Peng, Shen, Tu & Zhou leg.; SHNU • 2♂♂, 3♀♀; Mangshan N.R.; 10 May 2020; SHNU • 6♂♂, 12♀♀; Yanling County, Nanfengmian; 26°18'10"N, 114°00'12"E; alt. 1620 m; 26 May 2014; Peng, Shen, Yu & Yan leg.; SHNU • 1♂, 1♀; Yanling County, Nanfengmian; 26°16'32"N, 113°59'34"E; alt. 1380 m; 27 May 2014; Peng, Shen, Yu & Yan leg.; SHNU • 2♂♂, 1♀; Yanling County, Nanfengmian; 26°18'20"N, 114°00'51"E; alt. 1730 m; 28 May 2014; Peng, Shen, Yu & Yan leg.; SHNU. - Fujian Prov. • 15♂♂, 14♀♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1300-1500 m; 27 May 2012; Peng & Dai leg.; SHNU • 2♂♂, 1♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1200-1500 m; 26 May 2012; Peng & Dai leg.; SHNU • 2♂♂, 2♀♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1200-1500 m; 25 May 2012; Peng & Dai leg.; SHNU • 3♀♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1200-1300 m; 24 May 2012; Peng & Dai leg.; SHNU • 1♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1100-1400 m; 29 May 2012; Peng & Dai leg.; SHNU • 1♂, 2♀♀; Wuyishan City, Guadun Vill.; 27°44'N, 117°38'E; alt. 1800 m; 01 June 2012; Peng & Dai leg.; SHNU • 1♂; Guadun Vill.; August 2008; Zhu-Qing He leg.; SHNU • 2♂♂, 1♀; Guihe Vill., Meihua Mt.; alt. 1500 m; 20 May 2007; Huang & Xu leg.; SHNU • 1♂; N. Slope Gouzinao, Meihua Mt.; alt. 1650 m; 29 May 2007; Huang & Xu leg.; SHNU • 1♂; Guihe Vill., Gouzinao, Meihua Mt.; alt. 1500 m; 26 May 2007; Huang & Xu leg.; SHNU.

Measurements.

Male: BL: 9.0-10.8 mm, FL: 5.0-5.8 mm. HL: 1.22-1.56 mm, HW: 1.72-2.11 mm, CL: 0.89-1.06 mm, PO: 0.22-0.28 mm, PL: 1.78-2.00 mm, PW: 1.45-1.72 mm, EL: 2.11-2.45 mm, EW: 2.17-2.56 mm. HL/HW: 0.69-0.76, CL/PO: 3.20-4.00, PL/PW: 1.16-1.23, EL/EW: 0.95-0.97, HW/EW: 0.79-0.83, PW/EW: 0.66-0.67, HW/PW: 1.18-1.23. Female: BL: 8.2-11.7 mm, FL: 4.7-5.6 mm. HL: 1.22-1.45 mm, HW: 1.61-2.00 mm, CL: 0.83-0.89 mm, PO: 0.22-0.28 mm, PL: 1.67-2.00 mm, PW: 1.33-1.67 mm, EL: 2.00-2.56 mm, EW: 2.00-2.67 mm. HL/HW: 0.70-0.76, CL/PO: 3.00-3.75, PL/PW: 1.20-1.25, EL/EW: 0.95-1.00, HW/EW: 0.75-0.81, PW/EW: 0.63-0.68, HW/PW: 1.18-1.21.

Diagnosis.

The new subspecies differs from the nominate subspecies in the slightly shorter tempora, and entirely reddish to yellowish antennae and legs (except a dark band on the femora), while the nominate subspecies has almost entirely dark antennae, and black tibiae and tarsi. Even in paler (teneral) specimens of the nominate subspecies, the antennae and legs are at least partly darkened.

Distribution.

The subspecies is widespread in China (Zhejiang, Fujian, Hubei, Hunan, Sichuan, Guangxi, Guangdong, Anhui, Jiangxi).

Figures

Figures 46 - 52. Thoracostrongylus formosanus flavipes ssp. nov. 46, 47 habitus 48 - 51 aedeagus, ventral (48, 49) and lateral (50, 51) views 52 male abdominal sternite VIII. Scale bars: 1 mm (46, 47); 0.2 mm (48 - 52).

Figures 108 - 122. Distribution map of Thoracostrongylus species of China 108 T. acerosus 109 T. aduncatus 110 T. birmanus 111 T. diaoluoensis 112 T. formosanus formosanus 113 T. formosanus flavipes 114 T. fujianensis 115 T. malaisei 116 T. miyakei 117 T. sarawakensis 118 T. velutinus 119 T. baishanzuensis 120 T. bicolor 121 T. brachypterus 122 T. chrysites. Triangle, localities of specimens examined in this paper; square, localities of specimens listed in previous papers. Red, doubtful localities; blue, trustede localities.