Telothyria erythropyga Fleming & Wood, 2020

Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Dapkey, Tanya, Hallwachs, Winnie & Janzen, Daniel, 2020, Revision of Telothyria van der Wulp (Diptera: Tachinidae) and twenty-five new species from Area de Conservacion Guanacaste in northwestern Costa Rica with a key to Mesoamerican species, Biodiversity Data Journal 8, pp. 47157-47157 : 47157

Telothyria erythropyga Fleming & Wood sp. n.

Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007125 ; recordedBy: D.H. Janzen, W. Hallwachs & Anabelle Cordoba; individualID: DHJPAR0007125; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAV367-06, 06-SRNP-1595; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Puente Palma; verbatimElevation: 460; verbatimLatitude: 10.9163; verbatimLongitude: -85.3787; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9163; decimalLongitude: -85.3787; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen03; verbatimEventDate: 21-Mar-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011560 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011560; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ947-06, 05-SRNP-41630; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042; decimalLongitude: -85.2712; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 21-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011561 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011561; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ948-06, 05-SRNP-41628; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042; decimalLongitude: -85.2712; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 18-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011563 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011563; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ950-06, 05-SRNP-41977; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962; decimalLongitude: -85.2777; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Solis01; verbatimEventDate: 30-Aug-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011564 ; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona; individualID: DHJPAR0011564; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ951-06, 05-SRNP-42282; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962; decimalLongitude: -85.2777; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Desmia Janzen576; verbatimEventDate: 21-Sep-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011568 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011568; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ955-06, 04-SRNP-42880; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962; decimalLongitude: -85.2777; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 10-Jan-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0016623 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0016623; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAP827-07, 06-SRNP-44079; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Conguera; verbatimElevation: 420; verbatimLatitude: 10.9159; verbatimLongitude: -85.2663; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9159; decimalLongitude: -85.2663; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Desmia Solis19; verbatimEventDate: 25-Nov-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0018614 ; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0018614; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAI1261-07, 01-SRNP-23343; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962; decimalLongitude: -85.2777; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 05-Jan-2002; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0034439 ; recordedBy: D.H. Janzen, W. Hallwachs & Anabelle Cordoba; individualID: DHJPAR0034439; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYC1091-09, 09-SRNP-41069; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: San Lucas; verbatimElevation: 320; verbatimLatitude: 10.9185; verbatimLongitude: -85.3034; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9185; decimalLongitude: -85.3034; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 06-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035742 ; recordedBy: D.H. Janzen, W. Hallwachs & Anabelle Cordoba; individualID: DHJPAR0035742; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYD1123-09, 09-SRNP-41748; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Tucan; verbatimElevation: 410; verbatimLatitude: 10.9042; verbatimLongitude: -85.2712; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9042; decimalLongitude: -85.2712; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 14-Aug-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037318 ; recordedBy: D.H. Janzen, W. Hallwachs & Pablo Umana Calderon; individualID: DHJPAR0037318; individualCount: 1; sex: F; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYC4063-10, 09-SRNP-42882; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Rincon; verbatimElevation: 430; verbatimLatitude: 10.8962; verbatimLongitude: -85.2777; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8962; decimalLongitude: -85.2777; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Desmia Janzen07; verbatimEventDate: 10-Nov-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0039300 ; recordedBy: D.H. Janzen, W. Hallwachs & Elda Araya; individualID: DHJPAR0039300; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAV863-10, 10-SRNP-2485; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Tajo Angeles; verbatimElevation: 540; verbatimLatitude: 10.8647; verbatimLongitude: -85.4153; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8647; decimalLongitude: -85.4153; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Desmia Solis19; verbatimEventDate: 10-Jun-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0045686 ; recordedBy: D.H. Janzen, W. Hallwachs & Anabelle Cordoba; individualID: DHJPAR0045686; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ACGAZ875-11, 11-SRNP-43494; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Rio Francia Arriba; verbatimElevation: 400; verbatimLatitude: 10.8967; verbatimLongitude: -85.29; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8967; decimalLongitude: -85.29; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 28-Aug-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0051982 ; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0051982; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYH1094-13, 13-SRNP-70434; Taxon: scientificName: Telothyriaerythropyga; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Telothyria; specificEpithet: erythropyga; scientificNameAuthorship: Fleming & Wood, 2018; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Sendero Suspiro; verbatimElevation: 439; verbatimLatitude: 10.9839; verbatimLongitude: -85.3885; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9839; decimalLongitude: -85.3885; Identification: identifiedBy: AJ Fleming; dateIdentified: 2018; Event: samplingProtocol: Reared from the larva of the Crambidae, Phostria Janzen05; verbatimEventDate: 14-Apr-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Description

Male. Length: 5-9 mm (Fig. 22). Head (Fig. 22 b): frons narrow, 1/8 of head width; gena 1/9 of head height; four reclinate orbital setae; anteriormost reclinate orbital almost equal to uppermost frontal seta; ocellar setae present but so small as to appear absent; outer vertical seta absent; fronto-orbital plate brassy-gold on uppermost 30%; ocellar triangle with pale gold tomentum on posterior half, with anterior margin concolorous with frontal vitta; fronto-orbital plate with short blonde hairs interspersed among frontal setae; parafacial brilliant silver; facial ridge bare; palpus apically slightly inflated to appear slightly oar-shaped, sparsely haired along outer margin; arista brown, almost bare, distinctly-thickened on basal 1/10, microtrichia at most equally as long as width of arista; postpedicel orange over at most 60% of surface; postocular region behind margin of eye upper half gold, with lower half including gena silver tomentose; upper half of occiput gold tomentose. Thorax (Fig. 22 a, c): golden tomentose, with four distinct dorsal stripes; thorax entirely covered in dense plumose blonde hairs; chaetotaxy: 5-6 postpronotal setae, basal setae arranged in a straight line; supra-alar setae 2:3; intra-alar setae 1-2:2; dorsocentral setae 3:3; acrostichal setae 3:3; katepisternum with three setae. Scutellum golden tomentose; two pairs of strong marginal setae (basal and subapical) and a small pair of crossed apical scutellar setae 1/3rd as long as subapical scutellars; basal scutellar setae subequal in length to subapical setae; subapical setae straight; underside of scutellum bearing plumose blonde hairs below basal scutellar setae. Legs: foreleg with yellow ground color throughout, tibia and tarsal segments appearing darker due to hair covering; midleg, coxa and proximal half of femur yellow, remainder of femur, tibia and tarsal segments brown; hindleg dark brown extending from distal half of femur to tarsal segments; anterior leg tibia with regular fringe of equally spaced setae along anteroventral surface, and two strong posterodorsal seta. Wings: basicosta ivory white; all veins bare, with only 1-2 setulae at base of R4+5.; calypters pale white translucent with a narrow yellowish fringe. Abdomen (Fig. 22 a, c): ground color yellow-orange; ST1+2 brown over medial 30%, with yellow ventrolaterally, extending into a longitudinal middorsal brown stripe up to posterior edge of T3 and T4; T3-T5 with dense gold tomentum along anterior marginal 10%, thinning and extending over remainder of tergite; T5 entirely orange with gold tomentum; median marginal setae present only on T4 and T5; median discal setae absent. Male terminalia (Fig. 22 d, e, f): Sternite 5 with a wide deeply separated median cleft, widely V-shaped, margins tomentose; lateral lobes of sternite subtriangular apically, outer margins covered in strong setae; basal section of sternite 5 subequal in length to apical lobes. Cerci in posterior view sharply pointed rectangular, fused along entire length; medial shoulder weak to undeveloped, entire structure vaguely sword shaped. In lateral view cerci, with a mild downward curve, and several strong widely spaced setae along basal 2/3rds. Surstylus in lateral view rounded at tip, not downwardly curved, overall digitiform in appearance; fused with epandrium; when viewed dorsally surstyli appear straight with a very slight club or swelling apically. Distiphallus 2X as long as basiphallus and tubular, slightly pointed at apex.

Female. Length: 5-7 mm (Fig. 23). Head (Fig. 23 b): as in male with the following exceptions: fronto-orbital plate 30% gold; ocellar triangle concolorous with remainder of vertex; parafacial brilliant silver; frons 1/4 of head width; four inner reclinate orbital setae; two proclinate orbital setae; outer vertical seta present; palpus significantly inflated and oar-shaped apically. Thorax (Fig. 23 a, c): katepisternum with three setae; meron with a brush of plumose setae and 2-3 typical meral setae. Legs: colored as in male. Abdomen (Fig. 23 a, c): ground color brown dorsally on ST1+2 and T3 with orange laterally; T4 entirely brown ground color; T5 as in male; marginal setae present on T4, T5 and sometimes on T3.

Diagnosis

Telothyria erythropyga sp. n. can be distinguished from all other Telothyria by the following combination of traits: ocellar setae present but so small as to appear absent, parafacial brilliant silver, postpedicel orange over at most 60% of surface, arista plumose microtrichia at most equally as long as width of arista, facial ridge bare, thorax entirely covered in dense plumose blonde hairs, and T5 orange with gold tomentum.

Etymology

Telothyria erythropyga sp. n. From the Greek adjective " erythros " meaning red and the Greek noun " pygo " meaning rump or tail, in reference to the apically orange T5.

Distribution

Costa Rica, ACG, Alajuela and Guanacaste Provinces, 320-540 m elevation.

Ecology

Telothyria erythropyga sp. n. has been reared 33 times from four species of Lepidoptera in the family Crambidae : Phostria Janzen05, Phostria Janzen03, Desmia Solis19, Desmia Janzen576, Desmia Janzen07 in rain forest and dry-rain lowland intergrade.