Spinatrypina Rzhonsnitskaya, 1964

Ma, Xue-Ping, Becker, Ralph Thomas, Li, Hua & Sun, Yuan-Yuan, 2006, Early and Middle Frasnian brachiopod faunas and turnover on the South China shelf, Acta Palaeontologica Polonica 51 (4), pp. 789-812 : 802-803

Genus Spinatrypina Rzhonsnitskaya, 1964 Spinatrypina douvillei ( Mansuy, 1912)

Fig. 10A, B .

1912 Atrypa Douvillei nov. sp.; Mansuy 1912: 74, pl. 13: 9a–h.

1931 Atrypa douvillii Mansuy ; Grabau 1931: 194–197, pl. 15: 15, pl. 22: 5–10.

1938 Atrypa douvillii Mansuy ; Tien 1938: 95–97, pl. 13: 5–10, 13.

1983 Spinatrypina douvillii (Mansuy) ; Chen 1983: 316, pl. 31: 3–5.

Material.—About 50 poorly preserved shells, valves, and fragments from the Early Frasnian of the Caiziyan section (samples GC 8–9, GC 20, and GC 21–24). Three complete shells from the?Middle Frasnian of the Panxi section (sample PY5).

Discussion.—The specimen of Fig. 10A is a heavily exfoliated, weakly convex dorsal valve, with about 40–45 ribs (approximately 19.0 mm wide, 16.5 mm long, exact rib number is uncertain), with about 6 ribs per 5 mm near the anterior. Another figured specimen ( Fig. 10B ) is also a dorsal valve, which is subrounded in outline, slightly convex, with about (5–) 6 ribs per 5 mm at anterior margin (around 30–35 ribs on whole surface; size is about 17.0 mm wide, 13.9 mm long).

The outline, dorsal valve convexity, and nature of ribbing are basically in agreement with Spinatrypina douvillei ( Mansuy, 1912) and material described by subsequent workers from South China (e.g., Chen 1983) [see Grabau (1931: 194) for an English translation of Mansuy’s original French description]. Grabau (1931: 194) changed the name douvillei (to honor M.H. Douvillé by Mansuy 1912) to douvillii without giving any reasons, which has since been followed by subsequent workers, such as Tien (1938), Wang et al. (1964), and Chen (1983).

Occurrence.—Same as the previous species in geographic and stratigraphic distribution. The studied specimens are from the

Caiziyan and Panxi sections ( Fig. 2 ), correlated with the Palmatolepis transitans – Pa. punctata conodont zones.

Figures

Fig. 10. The spinatrypinae brachiopods from the Early and Middle Frasnian of the Caiziyan and Qiziqiao sections. A, B. Spinatrypina douvillei (Mansuy, 1912). A. PUM05018, sample GC21–24, Caiziyan section, Early Frasnian, dorsal view of an exfoliated specimen on slab. B. PUM05019, sample GC21–24, Caiziyan section, Early Frasnian, dorsal view of another exfoliated specimen on slab. C–G. Spinatrypa subkwangsiensis (Tien, 1938) Group B. C. PUM05020, sample QZQ1–3, Qiziqiao section, Middle Frasnian, dorsal view of an exfoliated specimen on slab. D. PUM05021, sample QZQ4, Qiziqiao section, probably Late Frasnian, dorsal (D1), lateral (D2), ventral (D3), posterior (D4), and anterior (D5) views of an adult(?), note spiny bases and broken spines. E. PUM05022, sample QZQ4, Qiziqiao section, probably Late Frasnian, posterior (E1), anterior (E2), ventral (E3), dorsal (E4), and lateral (E5) views of another adult (?) with a well preserved foramen bounded by a pair of conjunct deltidial plates at the bottom. F. PUM05023, sample QZQ4, Qiziqiao section, probably Late Frasnian, dorsal (F1), ventral (F2), lateral (F3), posterior (F4), anterior (F5), and enlarged area (F6) views of a juvenile with a well preserved foramen bounded by a pair of conjunct deltidial plates at the bottom. G. PUM05024, sample QZQ4, Qiziqiao section, probably Late Frasnian, anterior (G1), posterior (G2), dorsal (G3), ventral (G4), and lateral (G5) views of another juvenile. H–I. Spinatrypina lungkouchungensis (Tien, 1938). H. PUM05025, sample QZQ4, Qiziqiao section, probably Late Frasnian, ventral view, specimen is on slab. I. PUM05026, sample QZQ3, Qiziqiao section, Middle Frasnian, dorsal (I1), lateral (I2), ventral (I3), and anterior (I4) views of a juvenile.

Fig. 2. Stratigraphy and distribution of Early and Middle Frasnian brachiopods in South China. Most but not all known brachiopods are described and illustrated in this paper. For the distribution of atrypids in the Dushan section, for example, see Ma et al. (2005). The stratigraphy of Panxi and the occurrence of Cyrtospirifer sp. are based on Yunnan compiling group (1978) and the occurrence of Stringocephalus in the lower part of the Yidade Formation is as in Xian et al. (1988: 148). The stratigraphy of Dushan and the occurrence of Stringocephalus sp. are as in Liao et al. (1979); the Caiziyan section was re−measured by Long−Ming Wei, Xue−Ping Ma, and Hua Li (personal communication 2005), with coral identifications by Wei−Hua Liao. The stratigraphy of Qiziqiao is based on data of Z.−X. Hu and Z.−Q. Deng in Tan et al. (1987), and the occurrence of Yunnanella sp. is based on Zu−Han Liu (personal communication 2001). Abbreviations: calc., calcareous; argill., argillaceous; C., Cyrtospirifer; Pa., Palmatolepis; Po., Polygnathus; S., Spinatrypina.