Sidymella excavata Machado & Guzati

Machado, Miguel, Guzati, Catherine, Viecelli, Rafaela, Molina-Gomez, Diana & Augusto Teixeira, Renato, 2019, A taxonomic review of the crab spider genus Sidymella (Araneae, Thomisidae) in the Neotropics, Zoosystematics and Evolution 95 (2), pp. 319-344 : 319

Sidymella excavata Machado & Guzati sp. nov. Figures 2D, 3D, 4, 5

Type material.

Holotype ♀, ECUADOR: Cerro Troya, Carchi, 0°43'59.7"N, 77°41'00.3"W, L. Pena leg. (MCZ 133396). Paratypes: ♂, ECUADOR: Pichincha, 0°15'00.0"S, 78°35'00.0"W, 19.iv.1988, W. Maddison leg. (MCZ 133397); ♀, COLOMBIA: Cundinamarca, Mosquera ( Mondoñedo), 4°41'0.06"N, 74°15'0.25"W, 20.x.2000, E. Florez & J. Pinzon leg. (ICN 3404).

Material examined.

COLOMBIA: 2j, Magdalena (Santa Marta), 11°12'54"N, 74°06'01"W, 18.iv.1977, W. Galvis & A.J. Moreno leg. (ICN-Ar 9140); 1♀ 1j, Quindío ( Estación Bremen), 4°40'0"N, 75°39'0"W, 14-20.iv.1998, P. Ariza leg. (MPUJ 45511); 1♀, 1j, same data as preceding (MPUJ 11490); 1♀, Cundinamarca, Mosquera (via a La Mesa), 4°40'12.0"N, 74°16'00.7"W, 20.x.2000, E. Florez & J. Pinzon leg. (ICN-Ar 3404); 2♀, Mosquera (Desierto de Zabrinsky), 4°44'30"N, 74°14'8"W, 23.ii.2002, L. Benavides, C. Niño, A. Castañeda & G. Mora leg. (ICN-Ar 2157); 2♀, 5j, 06.iv.2002, same locality and collectors (ICN-Ar 2146); 4♀, 7j, same locality, April 2006, J. Martínez leg. (ICN-Ar 10500); 1♀, 23.ii.2002, same locality, C. Niño, G. Mora, A. Castañeda & J. Martínez leg. (CAUD-216-ARA 139); 1♀, 27.vii.2002, same locality, J. Martínez, C. Niño & G. Mora leg. (CAUD-216-ARA 366); 1♀, iv.2002, same locality, C. Niño, G. Mora, A. Castañeda & J. Martínez leg. (CAUD-216-ARA 1944); 1♀, 08.ix.2016, same locality, D. Molina leg. (ICN-Ar 10449); 1♀, same data as preceding (ICN-Ar 10498). 1♀, 2j, Boyacá, Villa de Levya, 5°40'21"N, 73°27'42"W, 09.vi.2001, L. Benavides leg. (ICN-Ar 1248). 1♂, Valle del Cauca, Saladito, 3°34'15.78"N, 76°36'9.27"W, W. Eberhard leg. (MCZ 133395). ECUADOR: 1♂, Quito, Pichincha, 0°15'00"S, 78°35'0"W, 19.iv.1988, W. Maddison leg. (MCZ 133397). 1♂, 2j, Azuay, Cerro Tinajillas, 3°10'60"S, 79°02'00"W, 18-21.iii.1965, L. Pena leg. (MCZ 133403). PERU: 1♂, Chavin, Ancash, 9°25'56"S, 77°32'07"W, 14.i.1972 (MACN-Ar 19068). ARGENTINA: 3♂, 4 j, Catamarca, Las Chacritas, 27°41'19.25"S, 65°55'24.97"W, 20.i.1987, P. Goloboff leg. (MACN-Ar 19098).

Etymology.

The specific name refers to the shape of the anterior border of the opisthosoma with a remarkable median re-entrance/excavation.

Diagnosis.

Females of S. excavata sp. nov. resemble those of S. marmorata sp. nov. by the large body size, short opisthosomal projections and long copulatory ducts; however, they can be distinguished by the presence of long needle-shaped macrosetae on the ocular area (Fig. 4B), and by having five mesial macrosetae on femora I and II (Fig. 4A). Males are similar to those of S. lucida , but can be recognized by cephalic setae and a basal branch of the RTA which is truncated instead of conical (Fig. 5 C–F).

Description.

Female: Prosoma, sternum and legs totally orange; legs I and II covered by numerous spiniform setae; opisthosoma predominantly light-yellow with a dark median stain on the dorsum that splits posteriorly towards to the tips of the projections (Fig. 4A). Epigynum with a wide septum formed by the median junction of the posterior folds of the epigynal plate (Fig. 4C, E); copulatory ducts hyaline, long and coiled, leading to a pair of walnut-shaped spermathecae (Fig. 4D, F).

Measurements: eyes sizes and interdistances: AME 0.03, ALE 0.09, PME 0.05, PLE 0.03, AME–AME 0.15, AME–ALE 0.09, PME–PME 0.17, PME–PLE 0.17, MOQ length 0.41, MOQ width 0.17; leg formula: 1 –2–4– 3: leg I - femur 3.40/ patella 1.36/ tibiae 3.04/ metatarsus 2.04/ tarsus 0.88/ total 10.72; II - 2.68/ 1.12/ 2.08/ 1.72/ 0.68/ 8.28; III - 1.40/ 0.72/ 0.88/ 0.80/ 0.60/ 4.44; IV - 1.76/ 0.76/ 1.12/ 0.96/ 0.60/ 5.20. Total body length 5.48; prosoma 2.48 length, 2.04 wide; opisthosoma length 3.00; clypeus 0.35 height; sternum 1.12 length, 1.04 width; endites 0.56 length, 0.20 width; labium 0.32 length, 0.36 width.

Male: Prosoma yellow with a darker median stain; legs I and II yellow with brownish spots on the femoral setiferous tubercles and at the distal portion of each leg segment; legs III and IV light-yellow. Opisthosoma as in female (Fig. 5A). Palpi with a well-developed pars pendula and embolus emerging at three o’clock, curling at the tip (Fig. 5 C–F); basal branch of RTA stout and directed retro laterally while the RTA is short, rounded and oriented vertically (Fig. 5D, F).

Measurements: eyes diameters and eyes interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.05, AME–AME 0.13, AME–ALE 0.05, PME–PME 0.11, PME–PLE 0.13, MOQ length 0.17, MOQ width 0.11; leg formula: 1 –2–4– 3: leg I - femur 2.04/ patella 0.72/ tibiae 1.92/ metatarsus 1.68/ tarsus 0.76/ total 7.12; II - 1.44/ 0.60/ 1.20/ 1.12/ 0.60/ 4.96; III - 0.72/ 0.60/ 0.56/ 0.40/ 0.40/ 2.68; IV - 0.96/ 0.36/ 0.60/ 0.52/ 0.40/ 2.84. Total body length 3.12; prosoma 1.56 length, 1.24 wide; opisthosoma length 1.56; clypeus 0.19 height; sternum 0.68 length, 0.62 width; endites 0.35 length, 0.13 width; labium 0.17 length, 0.25 width.

Distribution.

COLOMBIA: Magdalena, Quindío, Cundinamarca, Boyacá, and Valle del Cauca; ECUADOR: Pichincha and Azuay; PERU: Chavin; ARGENTINA: Catamarca (Fig. 15).