Senecio cocuyanus (Cuatrecasas) Cuatrecasas (1950: 43)

Salomón, Luciana, Sklenář, Petr & Freire, Susana E., 2018, Synopsis of Senecio series Culcitium (Asteraceae: Senecioneae, Senecioninae) in the Andean region of South America, Phytotaxa 340 (1), pp. 1-47 : 12

9. Senecio cocuyanus (Cuatrecasas) Cuatrecasas (1950: 43) . Culcitium cocuyanum Cuatrecasas (1940: 5) .

Type:— COLOMBIA. ‘Cordillera Oriental, Departamento de Boyacá: Sierra Nevada del Cocuy, o de Güicán, arenales en el valle de Las Lagunillas, 4200 m alt., 12 sept 1938, J. Cuatrecasas & H. García Barriga 1483 (holotype: COL 000005386 !; isotypes: F 0076805 F!, F 0076804 F!, US 00122834!). Fig. 7 .

Senecio santanderensis (Cuatrecasas) Cuatrecasas (1950: 45) . Culcitium santanderense Cuatrecasas (1942: 28 , tab. 3), syn. nov. Type:— COLOMBIA. Cordillera Oriental; Departamento de Santander: Páramo del Almorzadero, vertiente N, 3800 m alt., 28 Nov 1941, J. Cuatrecasas 13527 (holotype: COL 000005439!; isotypes: F 0049334F!, F 0076988F!, BC 634970!, BC 624141!, US 00122838!).

Distribution and Habitat ( Fig. 5 ): — Senecio cocuyanus is endemic to Colombia, in páramos at elevations of 3200– 4500 m.

Note: — Senecio santanderensis is here treated as a synonym of S. cocuyanus ; after having observed type materials and general collection for the species, we found specimens with ambiguous assignment to the species (e.g. Carlson 3 and Diaz-Piedrahita 99). Cuatrecasas distinguished both species by its indument (whiter and tenuous in S. santanderensis , yellowish white and denser in S. cocuyanus ), habit slender in S. santanderenis (vs robust in S. cocuyanus ), oblong-lanceolate leaves (vs. oblong-spathulate in S. cocuyanus ), shorter corollas (8–9 mm in S. santanderensis , 9–12 mm in S. cocuyanus ) and shorter pappus (8–9 mm vs. 10–12 S. cocuyanu s). These features are not constant and appear in different combinations in the specimens, making the species delimitation unclear.

Figures

FIGURE 5. Senecioser.Culcitium, distribution maps.A. Senecio alatopetiolatus (triangle); Senecio antisanae (star), Senecio betonicifolius (square); Senecio burkartii (circle); Senecio cajonensis (pentagon). B. Senecio campanulatus (star); Senecio candollei (square). Senecio canescens (circle); Senecio cocuyanus (triangle); Senecio culctioides (pentagon). C. Senecio cuencanus (star); Senecio doryphyllus (petagon); Senecio expansus (triangle); Senecio ferrugineus (square); Senecio gargantanus (circle). D. Senecio hypsobates (square); Senecio imbaburensis (star); Senecio involucratus (pentagon); Senecio josei (triangle); Senecio keshua (circle). E. Senecio leucophorbius (triangle); Senecio lingulatus (star); Senecio longepenicillatus (pentagon); Senecio mojandensis (square); Senecio nivalis (circle). F. Senecio otophorus (star); Senecio patens (square); Senecio pflanzii (triangle); Senecio piedrahitae (circle); Senecio pindilicensis (pentagon). G. Senecio prearoptorum (circle); Senecio punasessilis (triangle); Senecio puracensis (pentagon); Senecio hyoseridis (star); Senecio rhizomatus (square). H. Senecio serratifolius (pentagon); Senecio subinvolucratus (triangle); Senecio superandinus (star); Senecio superparamensis (square). I. Senecio supremus (triangle); Senecio tephrosioides (star); Senecio timidus (square); Senecio violifolius (*).

FIGURE 7. Senecio cocuyanus. A. Habit. B. Capitulum. C. Calycular bract. D. Phyllary. E. Floret. F. Stamen. G. Style branches.