Sedum oblanceolatum Clausen (1975: 404)

Zika, Peter F., Wilson, Barbara L., Brainerd, Richard E., Otting, Nick, Darington, Steven, Knaus, Brian J. & Nelson, Julie Kierstead, 2018, A review of Sedum section Gormania (Crassulaceae) in western North America, Phytotaxa 368 (1), pp. 448-450 : 448-450

9. Sedum oblanceolatum Clausen (1975: 404) . Figs. 1C , 9C , 35

Type: — UNITED STATES. Oregon: Jackson Co., road 0.8 km below Deadmans Point, ridge NW of Dutchman Peak, 1580 m, 8 July 1965, R. T . Clausen 651034 (holotype, BH; isotypes, BH, US).

The type sheets bear a second collection, of fruiting material, that are not part of the type material. The fruiting stems were gathered at the type locality on 7 August 1965, and were numbered Clausen 651034, as were the flowering shoots of the type.

Additional specimens examined: — UNITED STATES. CALIFORNIA. Siskiyou County: Applegate River drainage, N of Cook and Green Pass, 1370 m, 24 May 1987, Denton 4335 ( WTU) ; same site, 11 June 1991, Zika & Mumblo 11198 ( OSC) ; same site, 19 July 2011, Zika 25650 A ( CAS, HSC, RSA, UCR) ; Elliott Creek Road, 975 m, Rolle s.n. ( OSC) . OREGON. Jackson County: Tallowbox Lookout spur, 1310 m, 19 July 2012, Joachims & Busby OR 110-475 ( OSC) ; near Dutchman Peak , 1525 m, 13 July 1950, Whittaker 201 ( BH, WS) ; same site, 28 July 1977, Denton 4110 ( OSC, WTU) ; W of Little Red Mountain , 1500 m, 3 June 1992, Zika 11612 ( OSC) ; near Kinney Mountain , 1220 m, 15 June 1984, Rolle ( OSC) ; NW of Dutchman Peak , 1582 m, 3 July 1965, Clausen 651013 ( BH) , Clausen 651016 ( BH) ; same site, 18 July 2011, Zika 25636 & Lang ( GH, KNFY, UC, WTU) ; same site, 3 July 2014, Otting 3813 ( CAS, MO) .

Distribution and ecology: —Limited to southern Jackson County, Oregon, in the Siskiyous ecoregion ( Meyers et al. 2015), and the adjacent portion of Siskiyou County, California ( Fig. 22 ), in the Klamath Ranges ecoregion (Jepson eFlora 2017). Its habitat was sunny xeric rock outcrops and, less commonly, thin-soiled rocky slopes, occasionally in partial or full shade of open forest canopy, at elevations of 975–1740 m. Substrates were diverse, including diorite, granite, phyllite-schist, schist, metavolcanics, metasedimentary rocks, as well as ultramafics such as serpentine, soapstone, and peridotite.

Notes: — Sedum oblanceolatum ( Fig. 35 ) had oblanceolate rosette and stem leaves. The young leaves and inflorescences had a white granular and irregular waxy coating; the wax wore off with age. The inflorescence branches and sepals were also strongly glaucous, in addition to the granular wax, at least when young. The sepals were acute; the petals were white and relatively broad distally, acute, and ascending at anthesis. Fresh anthers were yellow. Similar S. albomarginatum differed in the color of its flowers ( Fig. 21 ), a coarser habit, separate follicles in fruit, and its allopatric range ( Fig. 22 ).

Hybrids: —Rare putative hybrids (Zika 25647B, 25650B), morphologically intermediate between two common putative parents at the site ( Sedum oblanceolatum, Zika 25650A; S. oregonense, Zika 25647A), were recorded at Copper Butte, Siskiyou Co., California ( Fig. 5C , Table 4).

Figures

FIGURE 1. Dense rosettes on vegetative shoots, without visible internodes between the leaves, were found in most species. A. Sedum flavidum, Trinity Co., California (Zika 26646 & Brainerd). B. Sedum laxum subsp. laxum, Del Norte Co., California (Zika 25927). C. Sedum oblanceolatum, Jackson Co., Oregon (Zika 25636 & Lang). D–E. Sedum rubiginosum, Tehama Co., California. D. Wild plant (Zika 26238). E. Cultivated plant (Wilson & Coberly CWG-10). F. Sedum sanhedrinum, Mendocino Co., California (Zika 26629 & Brainerd).

FIGURE 5. Stem leaves longer than wide, with a truncated base, were typical of most species of Sedum section Gormania. The stem leaves were commonly broadly obovate to elliptic or oblong, and were flattened or nearly cylindrical.A. Sedum citrinum, Del Norte Co., California (Zika 26185). B. Sedum laxum subsp. laxum, Del Norte Co. (Zika 25927). C. Sedum oblanceolatum × oregonense, Siskiyou Co., California (Zika 25647B). D. Sedum obtusatum, Nevada Co., California (Zika 26271). E–F. Sedum oregonense, Josephine Co., Oregon (Zika 25657). G. Sedum sanhedrinum, Lake Co., California (Zika 26639 & Brainerd).

FIGURE 9. The sepals were short compared to the length of fresh petals in most species. In all species the sepals elongated after anthesis, as the petals wilted. A–C. Relatively short sepals in early anthesis. A. Sedum citrinum, Del Norte Co., California (Zika 26193). B. Sedum oregonense, Jackson Co., Oregon (Zika 25666 & Wilson). C. Sepals elongating post-anthesis, Sedum oblanceolatum, Siskiyou Co., California (Zika 25650A). D–F. Relatively long sepals in early anthesis. D–E. Sedum paradisum subsp. paradisum. D. Shasta Co., California (Lindstrand III & Van Susteren NSR-8). E. Trinity Co., California (Zika 25920 & J.K. Nelson). F. Sedum paradisum subsp. subroseum, Sierra Co., California (Zika 26279).

FIGURE 21. Sedum albomarginatum. A. Habit. B. Two flowering stems, the oblanceolate rosette leaves and stem leaves were similar in length and outline, the stem leaves were consistently elongated proximally and distally. C. Rosette leaves, fresh ones waxy, older weathered ones green. D. Stem leaves, showing truncated base. E–G. Dense white granular wax coating on fresh leaves, bracts and sepals, easily rubbed off on fingers. F. Flower and bud, showing erect pale yellow petals, white granular wax coating was dense on fresh sepals and bracts, sparse on corolla. G. Densely white granular waxy rosette leaves. A–D, G. Plumas Co., California (Zika 25865). E–F. Butte Co., California (Zika 25867).

FIGURE 22. Distribution of Sedum albomarginatum, S. citrinum, S. marmorense, S. moranii, S. oblanceolatum, and S. patens, based on verified herbarium specimens.

FIGURE 35. Sedum oblanceolatum. A. Habit, dense rosettes, type locality, Jackson Co., Oregon (Zika 25636 & Lang). B–C. Rosette leaves and stem leaves similar, oblanceolate, Siskiyou Co., California (Zika 25650A). D. Stem leaves of a putative hybrid in a mixed population of S. oblanceolatum and S. oregonense, Siskiyou Co. (Zika 25649). Stem leaves are ascending and overlapping, as in S. oblanceolatum, but relatively broad, as in S. oregonense at the site. E. S. oblanceolatum, dense inflorescence, varying from capitate to cylindrical. F–H. Flowers and buds, showing ascending white petals with acute tips, yellow anthers, and granular-waxy young sepals. F, H. Zika 25650A. G. Zika 25636 & Lang.