Scydmaenus (Corbulifer) gushi, Jałoszyński, 2018

Jałoszyński, Paweł, 2018, ' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4472 (3), pp. 505-531 : 520-521

Scydmaenus (Corbulifer) gushi sp. n.

( Figs 36–38 , 60 , 70 )

Type material. HOLOTYPE: ( AUSTRALIA, NEW SOUTH WALES): ♂, two labels: " AUSTRALIA, NSW / Wild Cattle Creek / State Forest / 30°12'S, 152°45'E / 30 May 1993 " [white, printed], "Tom Gush / In leaf litter / Tom Gush / Collection 3659" [white, printed] ( ANIC). PARATYPES: 1 ♂, 1 ♀, same data as for holotype ( ANIC, cPJ).

Diagnosis. Male metatibia gradually and moderately broadened, slightly broader than metafemur, broadest slightly behind middle, mesal tibial margin bisinuate, lateral (external) tibial margin bisinuate with a deep concavity located clearly before middle, dorsolateral glandular 'pocket' present, with mesal margin bearing long setae; aedeagus in dorsal view broadest in basal fourth and nearly gradually narrowed distad, apex subtrapezoidal with rounded sides; apical projections very long and bent so strongly that their apices are directed proximad.

Description. Body of male ( Fig. 36 ) strongly convex, moderately dark brown, covered with setae slightly lighter than cuticle; BL 2.15–2.18 mm.

Head broadest at small eyes, HL 0.43–0.45 mm, HW 0.45–0.48 mm; tempora in lateral view as long as ~ 2 × length of eye, only slightly convergent posterad and posteriorly bent mesad at a slightly obtuse angle. Punctures on frons and vertex very fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL 1.08–1.13 mm, antennomeres I–VI each distinctly elongate, VII and VIII each transverse (VII indistinctly so), IX and X much larger than VIII, each slightly transverse, XI about as long as IX–X combined, nearly twice as long as broad.

Pronotum rounded, broadest slightly behind anterior third; PL 0.60 mm, PW 0.55–0.58 mm. Antebasal pits indistinct. Punctures on pronotal disc very fine and shallow, inconspicuous; setae short, sparse and suberect.

Elytra oval, broadest in front of middle; EL 1.13 mm, EW 0.90 mm, EI 1.25; apices separately rounded. Punctures small and superficial, barely noticeable; setae longer and denser than those on pronotum, suberect to erect.

Metatibia ( Fig. 60 ) in dorsal view gradually and moderately broadened, broadest slightly behind middle, mesal margin concave in proximal third, convex near middle and indistinctly concave in distal third, lateral (external) margin bisinuate with a deep triangular concavity located clearly before middle; dorsolateral 'pocket' present, with mesal margin bearing long setae.

Aedeagus ( Figs 37–38 ) moderately slender; AeL 0.48 mm; in dorsal view median lobe broadest near basal third, narrowing distad, very shallowly and broadly constricted in median region, and weakly narrowed in subapical region to form short subtrapezoidal apical portion with rounded sides; apical projections conspicuously long and strongly bent dorsoproximad at a sharp angle.

Female. Similar to male, but with unmodified metatibiae and much smaller eyes; length of temple equals to ~ 5 × eye in lateral view. BL 2.25 mm; HL 0.48 mm, HW 0.48 mm, AnL 1.10 mm; PL 0.63 mm, PW 0.60 mm; EL 1.15 mm, EW 0.93 mm, EI 1.24.

Distribution. New South Wales; south-eastern coast of Australia ( Fig. 70 ).

Etymology. This species is dedicated to Tim Gush, a collector of many interesting Scydmaeninae preserved at ANIC.

Remarks. Males of Scydmaenus gushi are most similar to those of S. kiwarrakianus ; these are the only species with moderately and gradually broadened metatibiae in which the deep concavity of the lateral (external) tibial margin is clearly before middle, so that the distal portion of tibia is distinctly longer than the proximal portion. These two species differ in the shape of the male metatibia; in S. gushi the lateral (external) margin bears only one concavity (vs. two in S. kiwarrakianus ). Also the aedeagi differ; in S. gushi the apical half of the median lobe is less narrowed distad and the apical projections are directed proximad (dorsad in S. kiwarrakianus ).

Figures

FIGURE 70. Distribution of Corbulifer species.

FIGURES 36–38. Scydmaenus (Corbulifer) gushi sp. n. Habitus of male (36), aedeagus in dorsal (37) and lateral (38) views.

FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65).