Scydmaenus (Corbulifer) gibraltar, Jałoszyński, 2018

Jałoszyński, Paweł, 2018, ' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4472 (3), pp. 505-531 : 521-523

Scydmaenus (Corbulifer) gibraltar sp. n.

( Figs 39–41 , 61 , 70 )

Type material. HOLOTYPE: ( AUSTRALIA, NEW SOUTH WALES): ♂, two labels: " NSW:NE: 10km of / GibraltarRange NP (HQ) / 29 Mar 1980 / G.B. Monteith " [white, printed], " QM Berlesate No. 213 / 29°36'S x 152°11'E / Rainforest / Sieved litter" [white, printed] (QM) . PARATYPES (8 exx.): NEW SOUTH WALES: 2 ♂♂, same data as for holotype ; 3 ♂♂, 3 ♀♀, " NSW:NE: Gibraltar / Range NP / 10 Nov 1980 / G.B. Monteith " [white, printed], " QM Berlesate No. 270 / 29°36'S x 152°11'E / Rainforest (Granite) / Sieved litter" [white, printed] (QM, cPJ).

Diagnosis. Male metatibia gradually and moderately broadened, indistinctly broader than metafemur, broadest slightly behind middle, mesal tibial margin bisinuate, lateral (external) tibial margin weakly convex in proximal half, strongly convex at middle and weakly concave in distal fourth, with a an angulate emargination nearly at middle, dorsolateral glandular 'pocket' present, with mesal margin bearing long setae; aedeagus in dorsal view broadest in basal fourth and nearly gradually narrowed distad, apical portion nearly parallel-sided; apical projections very long and bent dorsoproximad at a sharp angle.

Description. Body of male ( Fig. 39 ) strongly convex, moderately dark brown, covered with setae slightly lighter than cuticle; BL 1.88–1.95 mm (mean 1.90 mm).

Head broadest at small eyes, HL 0.35–0.40 mm (mean 0.38 mm), HW 0.38–0.43 mm (mean 0.40 mm); tempora in lateral view as long as 2.5 × length of eye, only slightly convergent posterad and posteriorly bent mesad at a slightly obtuse angle. Punctures on frons and vertex very fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL 0.95–1.03 mm (mean 0.99 mm), antennomeres I–VI each distinctly elongate, VII and VIII each slightly transverse, IX and X much larger than VIII, each distinctly transverse, XI about as long as IX–X combined, about twice as long as broad.

Pronotum rounded, broadest slightly behind anterior third; PL 0.50–0.55 mm (mean 0.53 mm), PW 0.45–0.50 mm (mean 0.48 mm). Antebasal pits indistinct. Punctures on pronotal disc very fine and shallow, inconspicuous; setae short, sparse and suberect.

Elytra oval, broadest in front of middle; EL 0.98–1.03 mm (mean 1.00 mm), EW 0.75–0.80 mm (mean 0.78 mm), EI 1.25–1.33; apices separately rounded. Punctures small and superficial, barely noticeable; setae longer and denser than those on pronotum, suberect to erect.

Metatibia ( Fig. 61 ) in dorsal view gradually and moderately broadened, broadest distinctly behind middle, mesal margin concave in proximal and distal thirds and convex in median third, lateral (external) margin weakly convex in proximal half, strongly convex at middle and indistinctly concave near distal fourth, with an angulate emargination indistinctly before middle; dorsolateral 'pocket' present, with mesal margin bearing long setae.

Aedeagus ( Figs 40–41 ) moderately slender; AeL 0.45 mm; in dorsal view median lobe broadest near basal third, strongly and gradually narrowing distad to form an elongate subtrapezoidal apical portion, with apical projections conspicuously long and strongly bent dorsoproximad at a sharp angle.

Female. Similar to male, but with unmodified metatibiae and much smaller eyes; length of temple equals 5–6 lengths of eye in lateral view. BL 1.95–2.00 mm (mean 1.98 mm); HL 0.38–0.40 mm (mean 0.39 mm), HW 0.40– 0.43 mm (mean 0.42 mm), AnL 0.95–0.98 mm (mean 0.96 mm); PL 0.55 mm, PW 0.48–0.50 mm (mean 0.48 mm); EL 1.05– 1.03 mm (mean 1.04 mm), EW 0.80 mm, EI 1.28–1.31.

Distribution. New South Wales; central-eastern coast of Australia ( Fig. 70 ).

Etymology. Locotypical, after the Gibraltar Range (a noun in apposition).

Remarks. Males of Scydmaenus gibraltar can be easily distinguished from similar congeners on the basis of the long distal portion of the metatibia which has its both lateral (mesal and external) margins weakly concave. Also the aedeagus of this species is unique in shape and easily distinguishable from the copulatory organs of all remaining Corbulifer spp.

Figures

FIGURE 70. Distribution of Corbulifer species.

FIGURES 39–41. Scydmaenus (Corbulifer) gibraltar sp. n. Habitus of male (39), aedeagus in dorsal (40) and lateral (41) views.

FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65).