Salcedia coquilhati Alluaud, 1932

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
publication LSID	lsid:zoobank.org:pub:843F6065-D00E-48E6-9A26-9CEB044BF102
persistent identifier	https://treatment.plazi.org/id/9A02A9FC-7D49-5695-BB7D-B3E4F23798EE
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia coquilhati Alluaud, 1932
status

Treatment

Salcedia coquilhati Alluaud, 1932 View in CoL Figs 11 View Figures 8–13 , 29 View Figures 26–35 , 55 View Figures 52–57 , 70 View Figures 68–76 , 84 View Figure 84

Salcedia coquilhati Alluaud, 1932: 3; Csiki 1933: 641; Alluaud 1935: 19; Burgeon 1935: 159; Reichardt 1975: 103; 1932, Dostal 1993: 121; Lorenz 2005: 155.

Type material.

Holotype: ♂, with labels and data: white, printed and handwritten “MUSÉE DU CONGO Coquilhatville –XII– 1924 DrH.Schouteden" / red, glued “TYPE” / white printed and handwritten "R. DÉT. 2019 f" / white, black framed, handwritten and printed " Salcedia coquilhati All. Alluaud det. 19" / red, black double framed “TYPE” / white, black printed "RMCA ENT 000019377" and square barcode (MRACT). Paratype: 1 ♀, with labels and data: white, printed and handwritten “MUSÉE DU CONGO Coquilhatville -XII- 1924 DrH.Schouteden" / white, handwritten " Salcedia coquilhati n.sp. Ch. Alluaud det." / red, black double framed “PARATYPE” (MRACT).

Remark.

In the holotype the left anterior leg is missing and the hind body shows some slight damages. In the paratype the left hind leg is missing.

Additional material.

1 ♂, 1 ♀, 1 spec., Eala I-1935 and XI-1934 J. Ghesquière MUSÉE DU CONGO; 1 ♂, 1 ♀, 2 specs., Irebu 30-XII-1920 DrH.Schouteden MUSÉE DU CONGO (MRACT, CBB); 10 specs., COLL. MUS. Congo Ubangi: Nouvelle-Anvers 9-XII-1952 P. Basilewsky / A la lumière (MRACT, CBB); 1 spec., I.R.S.A.-MUS.R.A.C. Equateur: terr. Bikoro, Mabali IX.1959 N. Leleup / Forêt inondée à la lumière (MRACT); 1 spec., Coquilhatville - a table le soir - G Juillet 1909 / MUSÉE DU CONGO Voyage de S.A.R. le Prince Albert 1909 (MRACT); 1 spec., MUSÉE DU CONGO Stanleyville a Coquilhatville Dr. Géranrd. (MRACT); 1 spec., MUSÉE DU CONGO Kai Bumba DrH.Schouteden 11-X-1920 (MRACT); 1 spec., COLL. MUS. CONGO Tshuapa: Bamania XII-1952 R. P. Hulstaert (MRACT); 1 spec., Congo Belge, P.N.G. Miss. H. De Saeger Mabanga 1949-1952 Réc. H. De Saeger (MRACT); 1 spec., Congo belge, P.N.G. Miss. H. De Saeger I/b/3 ’’, Mabanga 14-IV-1950 Réc. H. De Saeger.458 / Salcedia (MRACT); 1 spec., MUSEUM PARIS Afrique Équatoriale Fr. BRAZZAVILLE ANDRÉ GIDE 1925 / AOUT / Salcedia coquilhati Alluaud det. 1934 (MNHN).

Diagnosis.

A small sized species, with long-ovoid outline of the elytra with maximum width at posterior third. The pseudohumerus is obtuse angled and not dentate. The antennomeres are elongate. Distinguished from all other species of the genus by the extraordinary transverse pronotum which shows at the lateral margin five large transverse pits instead of six pits present in all other species, and the head exhibits a reniform outline in dorsal view.

Redescription.

Measurements in Table 1 View Table 1 .

Colour and surface: Melaneous, opaque; top of carinae griseous; mandibles and legs fuscous, antennae and palpi hinnuleous.

Head: Two-thirds of width of pronotum. Outline reniform. Clypeus wide, slightly but distinctly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by obtuse notches, frons with two flattened paramedian carinae, joining anteriorly into a convex bulge not clearly separated from frons, with two small glossy teeth bilaterally anterior to central bulge, with two parallel carinae paralaterally near base; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with moderately deep circular pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates distinctly margined, supra-antennal plates slightly vaulted, margin of supraorbital plates conspicuously raised. Base emarginated at middle, laterally rounded-off and constricted to neck without angle. Eyes large, convex, genae slightly convex, both of them clearly visible from above, with nearly circular shape in lateral view. Antenna with segments five to ten elongate (L/W 1.26), densely pubescent, segments two to four scarcely pubescent, scapus with minute longitudinal reticulation. Labrum straight anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, epilobes wide, projecting and nearly rectangular anteriorly, completely margined, surface coriaciate-like with flat impressions.

Pronotum (Fig. 29 View Figures 26–35 ): Outline rectangular, conspicuously transverse, nearly half times wider than long. Lateral margin straight but converging anteriorly, maximum width at posterior third, slightly rounded anteriorly and to posterior angles. Lateral margin distinctly crenulated, with nine tubercles, with distinct emargination at posterior angles. Base with wide tubercle laterally. Disc with two raised paramedian irregularly sinuate carinae parallel to median line, joining at anterior margin, with median line indistinct, with two additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly as less raised paralateral carina. With one distinctly long outer lateral carina running up to anterior margin. Lateral margin and space between lateral margin and lateral carina wing-like bent up, with five large transverse pits.

Elytron: Slightly convex in anterior half (lateral view), convex in frontal view. Outline in posterior five sixths long-ovoid, maximum width at middle. Pseudohumerus sub-rectangular, with small tubercle-like tooth. Apex acute at suture. Disc with interneur six crenulated, interneur two running up to apex as distinctly convex line, conspicuously raised; interneur four running in parallel to interneur six, reaching base, not reaching apex. Interneur four and six with two to three indistinct rows of pits becoming transverse apically.

Hind wings: Fully developed.

Lower surface: Pronotum with surface of antennal channel with irregular reticulation. Pseudoepipleura with two rows of serial pits, lateral margin of elytron smooth. Metepisternum elongate, with broad longitudinal groove. Metasternum and last five abdominal sternites with numerous pits, regularly arranged at carinae and margins. Sternum four to six distinctly sulcate.

Legs: Profemora with dorsal surface indistinctly reticulated. Protibia with terminal spine arcuate at apex, laterally with three teeth, dorsally and ventrally with two carinae. Movable spur arcuate, acute, length nearly as half as first tarsomere. First tarsomeres distinctly elongated, almost as long as tarsomeres two to five together.

External sexual dimorphism: Not observed.

Male genitalia (Fig. 55 View Figures 52–57 ): Median lobe short, stout, in dorsal view straight, in lateral view straight, with a few fine pili in apical third, oroficium small, apex circular in cross-section, slightly turned dorsally. Endophallus with two elongated groups of microtrichia, groups as long as one fifth of median lobe. Dorsal paramere sinuate in apical half, lateral and basal apophysis slender; both parameres slightly distorted.

Female genitalia (Fig. 70 View Figures 68–76 ): Coxostylus less sclerotised, broadened just anterior base, distinctly curved dorsally and slightly medially, slender in apical third, at end of basal third with one shorter strong and three slender nematiform setae laterally, with another slender seta at middle, with one long strong nematiform seta ventrally.

Variation: On the elytron between the interneurs four and six, the rows of pits vary in the number of pits and rows. In some of the specimens the labrum is indistinctly convex.

Distribution

(Fig. 84 View Figure 84 ). Known from several localities along the course of the Congo River and at the mouth of the river Tshuapa. Some of the specimens were collected at light. The historic name Coquilhatville refers to the city today called Mbandaka, that of Stanleyville to Kisangani, and Brazzaville to Kinshasa.