Rasnitsyneura aequabilis ( Bode, 1953 ) Makarkin & Ansorge & Khramov, 2021

Makarkin, Vladimir N., Ansorge, Jörg & Khramov, Alexander V., 2021, Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera), Palaeoentomology 4 (6), pp. 516-531 : 521-522

Rasnitsyneura aequabilis ( Bode, 1953) , comb. nov.

( Figs 5–7 )

Prohemerobius aequabilis Bode, 1953: 252 , 257, 258; Pl. 12, Fig. 326; Ponomarenko, 1996: 75, 76, Figs 2 , 10 (as a synonym of Prohemerobius septemvirgatus Bode, 1953 ).

Prohemerobius aequalibis [sic]: Ponomarenko, 1996: caption to Fig. 10 .

Material. Specimen PIN 5552/34 (part only); a wellpreserved, nearly complete forewing; deposited in PIN.

Locality and horizon. Germany: Schandelah. Early Jurassic (early Toarcian, boreale nodule, Harpoceras falciferum ammonite zone).

Description. Specimen PIN 5552/34 ( Fig. 5 ). Forewing 7.8 mm long, 2.6 mm wide. Costal margin medially markedly concave. Costal space relatively narrow, broadened proximally, distally. Humeral veinlet recurrent, with two short branches; terminal parts of proximal subcostal veinlets strongly bent toward wing apex, simple or with very shallow terminal fork; middle veinlets terminally curved to apex or forked with distal branch longer than proximal; distal veinlets of Sc, RA mainly once forked (two veinlets forked two or three times). Subcostal space moderately broad distally; only basal crossvein detected located slightly distad origin of RP. Sc, RA stout, distally approaching, probably not fused. RA space narrow, slightly dilated basally; five, rather regularly spaced crossveins in middle part. RP originating near wing base; anterior trace forked distally, not zigzagged, with seven branches. RP1, RP2, RP4 dichotomously forked distad outer gradate series; RP3, RP5, RP7 once shallowly forked; RP6 dichotomously forked far proximad outer gradate series. One short crossvein between R, Min basal part (1r-m) detected, connecting R, M. Fork of Mnot preserved. Anterior trace of MA forked distally, with one relatively short branch. MP pectinately forked, with five long branches, once to twice shallowly forked (one branch simple). Crossveins in radial to medial spaces poorly preserved; four crossveins (from RP2 to RP6) represented outer gradate series. Cu dividing into CuA, CuP relatively near wing base. CuA pectinately forked, with two, very long branches; each branch once shallowly forked. CuP shallowly once forked. One (proximal) crossvein between CuA, CuP detected. Two crossveins between CuP, A1: basal crossvein short; distal crossvein rather long. A1 profusely, pectinately branched, with two rather long branches, which are again pectinately or dichotomously branched. A2 pectinately forked, with two short branches. Two long crossveins between A1, A2. A3 fragmentarily preserved. Maculation: eight brown spots medially (different in size), two paler spots apically.

Remarks. We assign this specimen to Rasnitsyneura aequabilis judging from the drawings of the holotype by Bode (1953: Fig. 326) and Ponomarenko (1996: Fig. 10 , which we consider mostly correct). Both specimens have relatively narrow forewings, almost identical maculation shown by Bode (1953), and similar venation as shown by Ponomarenko (1996) if the anterior branch of Mis interpreted as RP1, the posterior branch of Mas MA, CuA as MP, and CuP as CuA. The holotype of R. aequabilis is also from Schandelah.

Figures

FIGURE 2. Polyosmylus excelsus Ansorge, 1996, holotype LGA 1033. A, Photograph. B, Forewing venation. C, Hind wing venation. Scale bar = 1 mm (all to same scale).

FIGURE 5. Rasnitsyneura aequabilis (Bode, 1953), specimen PIN 5552/34. A, Photograph. B, Forewing venation. Scale bar = 1 mm (both to same scale).

FIGURE 6. Rasnitsyneura sp., forewings. A and B, PIN 5552/36. C and D, PIN 5552/39. E and F, PIN 5552/38. G and H, PIN 5552/40. Scale bars = 1 mm (A and B, C–H to same scale).

FIGURE 7. Rasnitsyneura sp., specimen PIN 5552/37. A, Photograph. B, Forewing venation. Scale bar = 1 mm (both to same scale).

FIGURE 10. The hind wings of the Jurassic Polystoechotidae. A, An undescribed species of Polystoechotidae from the Early Jurassic (Sinemurian/ Hettangian) of Sogyuty (Kyrgyzstan), incorrectly assigned to Mesopolystoechus apicalis Martynov, 1937 by Martynova (1949: Fig. 8), PIN 371/323. B, The holotype of Jurapolystoechotes melonolomus Ren et al., 2002 from the late Middle Jurassic / early Late Jurassic of Daohugou (China), CNU-NEU-NN99021. Scale bars = 5 mm.