Pseudojulus carolinensis Shelley, 2004

Shelley, Rowland M., 2007, Rediagnoses of the milliped genera Pseudojulus Bollman, 1887, and Arvechambus Causey, 1963, in the southeastern USA; description of P. mississippiensis, n. sp. and proposal of the subtribe Pseudojulina (Julida: Parajulidae: Parajulinae: Aniulini), Zootaxa 1541, pp. 1-16 : 10

Pseudojulus carolinensis Shelley, 2004

Figs. 11–13

Pseudojulus carolinensis Shelley, 2004:198 –200, figs. 20–25.

Type specimens. ɗ holotype & Ψ paratype ( NCSM) and adult & immature ɗ paratypes ( SMNH) collected by D. Flynn, 6.5 mi (10.4 km) S Gastonia, along SR 1108 ca. 1.2 mi (1.9 km) SW jct US hwy. 321 and ca. 1.2 mi (1.9 km) N of South Carolina state line, Gaston Co., North Carolina. The holotype and adult ɗ paratype were collected on 21 May 1999, the juvenile ɗ paratype on 24 May 1999, and the Ψ paratype on 1 June 1999 ( Shelley 2004).

Diagnosis. A moderate-sized species characterized by basal fusion of otherwise distinct ag coxal lobes and distinct, widely segregated posterior syncoxal processes; pg telopodite crescent-shaped, angling caudad then curving broadly anteriad, with small but noticeable subapical lobe on inner/dorsal margin, prefemoral process extending caudal to telopodite basally then angling across curvature of latter. Cyphopodal gynaspis short, lobes indistinct.

Va r ia ti o n. The male from Jenkins Co., Georgia (figs. 11–13), within the distribution of P. paynei , displays features of this species plus P. carolinensis and coastalis and may represent an intergrade between all three species or just the last two, with whom it shares more attributes. With only one male and no material from central South Carolina, it cannot be unequivocally assigned, so I place it under P. carolinensis , which it best conforms to. On the ags, the posterior syncoxal lobes are laminate and clearly segregated, as in both P. c a ro - linensis and coastalis , and not fused, knob-like structures as in P. paynei ; however, the coxal lobes are low, basally fused, triangular structures, as in both P. paynei and carolinensis , and not the elongated, widely segregated projections of P. coastalis . The configuration of the pg is incompatible with P. paynei in that both the telopodite and prefemoral process curve strongly anteriad in a parallel arrangement. The prefemoral process lies over the medial surface of the telopodite and narrows and widens as if intermediate between the narrow projection in P. carolinensis and the wide, blade-like one in P. coastalis . The telopodite expands apically and possesses a prominent, subapical spur on the inner/dorsal surface that appears to be homologous to that on the stem in P. c o a s t a l i s, though located farther distad.

Distribution. The only published record is the type locality, although P. c a ro l i n e n s i s surely occurs in at least York & Cherokee cos., the adjacent cos. in South Carolina. The present new record, some 185 mi (296 km) south of the type locality, spans the entire state of South Carolina where P. c a ro l i n e n s i s was predicted to occur by Shelley (2004) it is also around 90 mi (144 km) west of the type locality of P. coastalis .

GEORGIA: Jenkins Co., Millen, ɗ, Ψ, 16 April 1940, collector unknown (FSCA).

Figures

FIGURES 11 – 13. Pseudojulus carolinensis. 11, right anterior gonopod of male from Jenkins Co., Georgia, anterior view. 12, the same, caudal view. 13, left posterior gonopod of the same, medial view. Scale line = 0.63 mm for fig. 23, 1.00 mm for figs. 21 – 22.