Pristiograptus lodenicensis Přibyl, 1943

Urbanek, Adam, Radzevičius, Sigitas, Kozłowska, Anna & Teller, Lech, 2012, Phyletic evolution and iterative speciation in the persistent Pristiograptus dubius lineage, Acta Palaeontologica Polonica 57 (3), pp. 589-611 : 606

Pristiograptus lodenicensis Přibyl, 1943

Figs. 4B, 5H, 6G, 10 .

1943 Pristiograptus lodenicensis sp. nov.; Přibyl 1943: 23–24, textfigs. C, D, J, K; pl. 3: 8–11.

1952 Pristiograptus lodenicensis Přibyl, 1943 ; Münch 1952: 91, pl. 20: 7.

1967 Pristiograptus lodenicensis Přibyl, 1943 ; Gailite et al. 1967: 244, text−fig. 58; pl. 29: 4, 5.

1974 Pristiograptus lodenicensis Přibyl, 1943 ; Ulst 1974: 115, pl. 13: 1, 2a, b, pl. 11: 7.

2000 Pristiograptus lodenicensis Přibyl, 1943 ; Radzevičius and Paškevičius 2000: 97–98, pl. 4: 1a–c, 2a–c, pl. 6D.

2003 Pristiograptus lodenicensis Přibyl, 1943 ; Radzevičius 2003: 10– 11, figs. 11A–H, 12A–F.

2005 Pristiograptus lodenicensis Přibyl, 1943 ; Radzevičius and Paškevičius 2005: 165, pl. 2: F.

2006 Pristiograptus lodenicensis Přibyl, 1943 ; Radzevičius 2006: 337, fig. 4C–F.

Remarks.—The thecae are slightly curved, and widen towards their apertures. The ss angle is acute, and the λ angle is right. The thecal lips are symmetrically curved ( Fig. 9A, E ). The depressions and curvature of lips are variable in proximal and distal thecae. The lips maybe slightly or much curved, they may widen towards their ends, and may have small or large ventral incisions and curvatures close to their ends. In some specimens there are large symmetrical “stingray wing” structures located at the ends of lips ( Fig. 9F ). These structures may occur in a few thecae or all the thecae ( Radzevičius 2006: fig. 4c–f). The thecal apertures are right angles to the rhabdosome axis. Similar thecae occur in P. d. labiatus, thus both forms are not typical members of P. dubius group. This type of theca is similar to those of Istrograptus Tsegelniuk, 1976 and Neocolonograptus Urbanek, 1997 . This species is separated from the lodenicencis group ( Radzevičius and Paškevičius 2000).

P. praelodenicensis is very similar to P. lodenicensis with thecae slightly curved, widening towards the aperture, which shows a small ventral depression. Thecae of P. praelodenicensis do not have such kind of depression.

Stratigraphic and geographic range.—Upper Wenlock Cyrtograptus lundgreni Biozone ; Latvia, Lithuania, Poland, Czech Republic.

Figures

Fig. 10. SEM micrographs of two Homerian (Wenlock) graptolite Pristiograptus forms. A, B. Pristiograptus dubius paezerensis subsp. nov., mature rhabdosomes; Cyrtograptus lundgreni Biozone. A. ZPAL G.44/17, Bartoszyce IG−1 borehole, Poland, depth 1663.2 m. B. ZPAL G.44/19, Zawada borehole, Poland, depth 1555.9–1562 m. C, D. Pristiograptus dubius parvus Ulst, 1974 growing rhabdosomes, Bartoszyce IG−1 borehole, Poland, depth 1660.7 m; Pristiograptus parvus Biozone. C. ZPAL G.44/25. D. ZPAL G.44/26.

Fig. 9. SEM micrographs of the graptolite Pristiograptus lodenicensis Přibyl, 1943; Cyrtograptus lundgreni Biozone, Lower Homerian, Wenlock. A–D, F. Zawada borehole, Poland. A–C. Depth 1546.5–1552.7 m. A. ZPAL G. 44/13, general view of rhabdosome (A), lips of th3 and th4 (A), and th1 1 2 with sicula (A3). B. ZPAL G.44/14, general view of dorsal side of rhabdosome (B1), proximal end (B2). C. ZPAL G.44/15, general view of rhabdosome. D, F. Depth 1540–1546.5 m. D. ZPAL G.44/16, proximal end of rhabdosome. F. ZPAL G.44/17, proximal end of rhabdosome (F) and th2 lip with cortical 1 tissue “script” (“manta ray wing”) (F2). E. VU P.P9−4b, Parovėja−9 borehole, Lithuania, depth 561.9 m, general view of rhabdosome (E1), depressions of thecal apertural lip (E2).