Phylloicus quitacalzon, Prather, 2003

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 92-94

Phylloicus quitacalzon , new species

Figs. 97 , 98

Phylloicus quitacalzon is very similar to P. hansoni . It is distinguished by single­lobed mesal coremata, and the short truncate posterior process of tergum IV ( Fig. 97F ).

Adult. Forewing length 10.7­12.2 mm, n = 26.

Head chestnut brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden brown. Legs golden brown. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, golden; golden setae marking wing­coupling setae proximally. Hind wing basal brush present in male.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired posterior processes, mesal coremata and lateral coremata; posterior process short, rounded; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata single­lobed, without setae. Tergum V without sclerotized modifications ( Fig. 97F ). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched, notch deep and round; with small sclerite in intersegmental membrane posterolateral to sternite ( Fig. 97A, C ). Tergum IX with paired mesal ridges extending 1/2 length of segment; posterior margin smoothly rounded ( Fig. 97B ); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae absent ( Fig. 97A ). Preanal appendage longer than tergum X, but less than 11/2 times length, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage ( Fig. 97A, B ). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes long, length at least twice diameter; apex, in lateral view, rounded, in dorsal view, notched, notch shallow ( Fig. 97A, B ). Harpago rounded; peglike setae many, apical ( Fig. 97A, C ). Phallic endotheca with paired basolateral lobes, basolateral lobes large and round; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 97D, E ).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII ( Fig. 98C ). Tergum IX without mesal ridge; posterior margin marked mesally by small patch of thin cuticle ( Fig. 98B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 98A ). Tergum X appendage shorter than mesal lobe, base indistinct, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes marked only by slightly raised thin cuticle, or absent ( Fig. 98B ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 98A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite ovoid; posterior end of spermatheca a sclerotized ovoid ( Fig. 98A ).

Holotype male: PERU: Madre de Dios: Toma del Agua, Amazonia Lodge , 12°52'13”S, 71°22'34"W, 415 m, 29.vi.1993, Blahnik & Pescador ( MHNJP).

Paratypes: PERU: Cuzco: Paucartambo to Pilcopata rd., Quebrada Quitacalzón at Puente Quitacalzón , 13°01'34”S, 71°29'58"W, 1050 m, 25­27.vi.1993, Blahnik & Pescador — 1 female, 1 male ( MHNJP) ; — 2 females, 1 male ( NMNH) ; — 1 female, 1 male ( UMSP) ; Quincemil ,. xi.1962, Peña G. — 1 female, 1 male ( CNC) ; — 1 male ( UMSP) ; Paucartambo, Paucartambo to Pilcopata rd. , streamlet 50 m E Quinta Calzon, 1050 m, 26.vi.1993, Blahnik & Pescador — 1 male ( MHNJP) ; — 2 males ( NMNH) ; Quinta Calzon ca. 30 km NW Pilcopata, km 164, streamlet, 50 km E Quinta Calzon , 13°09'00”S, 71°22'00"W, 1030 m, 2.ix.1989, Adams — 1 male ( NMNH) ; Huanuco: Tingo Maria , 672 m, 25­31.i.1980, Heppner — 1 female ( NMNH) ; 1­6.ii.1980, Heppner — 4 females, 5 males ( NMNH) , Madre de Dios: Toma del Agua, Amazonia Lodge , 12°52'13”S, 71°22'34"W, 415 m, 29.vi.1993, Blahnik & Pescador — 1 male ( NMNH) .

Distribution. Peru.

Etymology. Named for the type locality.

Figures

FIGURE 97. Phylloicus quitacalzon. Male (A­C, UMSP000068290; D­E, UMSP000063340; F, holotype): A—lateral view; B—dorsal view; C—sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F tergum IV, dorsal view.

FIGURE 98. Phylloicus quitacalzon. Female (UMSP000063342): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view.