Philonthus turneri, Cameron, 1951

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2001 (265), pp. 2441-3020 : 2441-3020

turneri Cameron, 1951 , see: pullatus Herman, 2001 .

ulpius Smetana, 1995e: 653 ( Gabrius ; Type locality: Quebec: Rigaud. Also cited from

Nova Scotia, Ontario, Quebec, Georgia, Michigan, New Hampshire, New York, North Carolina, Ohio, West Virginia).

- DISTRIBUTION: Canada, USA.

ultimus Schillhammer, 1997: 12 ( Gabrius ; Type locality: Nepal: [ Patan District ], Kathmandu District, Phulcoki, 2600 m).

— Schillhammer, 2000 c: 119 ( Gabrius ; Nepal).

- DISTRIBUTION: Nepal.

unculus Tottenham, 1956b: 222 ( Gabrius ; Type locality: S. Rhodesia: Mashonaland, Salisbury ).

- DISTRIBUTION: Zimbabwe.

unzenensis Bernhauer, 1939e: 153 ( Philonthus ; subgenus Gabrius ; Type locality: Japan: Unzen bei Shimabara).

— Smetana, 1973b: 129 ( Gabrius ; characters; notes).

— Smetana, 1984: 125 ( Gabrius ; characters; Japan).

— Cho and Lee, 1997: 272 ( Gabrius ; characters; Korea).

- DISTRIBUTION: Japan.

ussuriensis Schillhammer, 1991: 53 ( Gabrius ; Type locality: USSR: Nikolsk Ussurijsk, Ussurigebirge).

- DISTRIBUTION: Russia.

velox Sharp, 1910: 130 ( Gabrius ; Type locality: Anglia: ad flumen Lymington).

— Bernhauer and Schubert, 1914: 359 ( Philonthus ; catalog).

— Gridelli, 1920: 140 ( Gabrius ; characters; Englnad).

— Scheerpeltz, 1933: 1367 ( Philonthus ; catalog).

— Wüsthoff, 1934: 61 ( Philonthus ; illustration of aedeagus).

— Wüsthoff, 1937: 32 ( Philonthus ; illustration of aedeagus).

— Fagel, 1945: 41 ( Gabrius ; characters; Belgium).

— Coiffait, 1951: 111, 115 ( Gabrius ; characters in key; France).

— V. Hansen, 1952: 90 ( Philonthus ; subgenus Gabrius ; characters; Denmark).

— Smetana, 1952b: 123 ( Gabrius ; characters; Czechoslovakia).

— Smetana, 1957: 65, 69 ( Gabrius ; characters; England; France; Belgium; Holland; Germany; Czechoslovakia; Austria; Scandinavia).

— Smetana, 1958: 251 ( Gabrius ; characters; ecology; Czechoslovakia).

— Smetana, 1960c: 333 ( Gabrius ; characters; England; Germany; Czechoslovakia; Austria).

— Palm, 1963b: 106 ( Philonthus ; subgenus Gabrius ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 190 ( Gabrius ; characters; central Europe).

— Horion, 1965: 192 ( Gabrius ; Britain; Norway; Sweden; Finland; Russia; Denmark; Holland; Belgium; France; Czechoslovakia; Hungary; Austria; Germany).

— Smetana, 1966: 331 ( Gabrius ; Czechoslovakia).

— Coiffait, 1974: 60 ( Gabrius ; characters; Europe; Britain; France).

— Pope, 1977: 30 ( Gabrius ; Britain).

— Tóth, 1984: 89 ( Gabrius ; characters).

— Lucht, 1987: 106 ( Gabrius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Lohse and Lucht, 1989: 169 ( Gabrius ; note).

— Smetana, 1993b: 49 ( Gabrius ; Czech Republic; Slovakia).

— Smetana, 1995e: 626 ( Gabrius ; characters; notes; Maryland).

- DISTRIBUTION: Europe, USA.

austriacus Scheerpeltz, 1947: 283 ( Gabrius ; Type locality: Österreich: Marchfeld , Niederösterreich).

— Smetana, 1952b: 124 ( Gabrius ; characters).

— Smetana, 1957: 65, 69 ( Gabrius ; characters; Austria).

— Smetana, 1960c: 335 ( Gabrius ; characters; Austria).

— Lohse, 1964: 190 ( Gabrius ; characters; central Europe).

— Horion, 1965: 193 ( Gabrius ; Austria).

— Coiffait, 1974: 59 ( Gabrius ; characters; Austria).

— Tóth, 1984: 89 ( Gabrius ; characters).

— Lucht, 1987: 106 ( Gabrius ; checklist; Austria).

— Lohse and Lucht, 1989: 169 ( Gabrius ; synonym of velox ).

— Smetana, 1995e: 626 ( Gabrius ; synonym of velox ).

ventralis Stephens, 1833: 252 ( Gabrius ; [Note: Nomen dubium]; Type locality: Suffolk, and on the shores of the Severn , near Bristol).

— Stephens, 1839: 400 ( Gabrius ; characters; England).

- DISTRIBUTION: Britain.

vernalis Gravenhorst, 1806 , see: osseticus Kolenati, 1846 .

vibius Smetana, 1995e: 681 ( Gabrius ; Type locality: Quebec: Rigaud. Also cited from

New Hampshire, New York, Ohio, Pennsylvania, Vermont, West Virginia).

- DISTRIBUTION: Canada, USA.

vicinus Cameron, 1932: 149 ( Philonthus ; Type locality: Simla Hills: Kotgarh, alt. 7000 feet; [Note: See lectotype designation by Schillhammer, 1997]).

— Scheerpeltz, 1933: 1367 ( Philonthus ; catalog).

— Schillhammer, 1997: 47 ( Gabrius ; lectotype designation; characters; India; Nepal; Pakistan) .

- DISTRIBUTION: Pakistan, India, Nepal.

viduus Cameron, 1933b: 346 ( Philonthus ; subgenus Gabrius ; [preoccupied]; Type locality: British North Borneo, Mt. Kinabalu, Kamborangah, 7200 feet; [Note: See lectotype designation by Schillhammer, 1997]. Also cited from Lumu Lumu, 5500 feet).

— Cameron, 1937: 6 ( Philonthus ; Java).

— Hammond, 1984: 194, 195 ( Gabrius ; Borneo).

— Schillhammer, 1997: 63 ( Gabrius ; lectotype designation: British North Borneo, Mt. Kinabalu, Kamborangah, 7200 feet; characters; Malaysia) .

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Borneo, Malaysia, Indonesia.

villosulus Stephens, 1833 , see: Neobisnius .

vindex Smetana, 1995e: 648 ( Gabrius ; Type locality: Quebec: Gatineau Pk., Blind Lake . Also cited from Manitoba, Alaska, and Minnesota).

— Smetana, 1965: 29 ( Gabrius ; [Note: Misidentified as seattlensis according to Smetana, 1995e: 648]; characters; Newfoundland).

- DISTRIBUTION: Canada, USA.

virilis Horn, 1884: 215 ( Philonthus ; Type locality: Vancouver; [Note: See lectotype designation by Smetana, 1995e]).

— Bernhauer and Schubert, 1914: 361 ( Philonthus ; catalog).

— Hatch, 1957: 200 ( Philonthus ; subgenus Gabrius ; characters; British Columbia; Washington; Oregon).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 705 ( Gabrius ; lectotype designation; characters; notes; British Columbia; Oregon; Washington) .

- DISTRIBUTION: Canada, USA.

visokeanus Cameron, 1950 g: 39 ( Philonthus ; subgenus Gabrius ; Type locality: Belgian Congo: Visoke, alt. 2800–3300 m).

— Schillhammer, 1997: 83 ( Gabrius ).

- DISTRIBUTION: Congo.

vulcanicola Schillhammer, 1997: 65 ( Gabrius ; Type locality: Java: W Java, Mt. Gede , 50 km

SE Bogor, 2600 m).

vulcanus Cameron, 1950 g: 40 ( Philonthus ; subgenus Gabrius ; Type locality: Belgian Congo: Shamuheru, volc. Nyamuragira, alt. 1843 m).

— Tottenham, 1956b: 219 ( Gabrius ; characters).

- DISTRIBUTION: Congo.

wawawai Hatch, 1957, see: punctatellus Horn, 1884 .

xanthipes Schillhammer, 1997: 43 ( Gabrius ; Type locality: Nepal: [Sindhupalchok District], Prov. Bagmati, Gul Bhanjyang , 2600 m).

- DISTRIBUTION: Nepal.

yakinius Tottenham, 1956b: 210 ( Gabrius ; Type locality: Cape Colony: Albany dist. ).

- DISTRIBUTION: South Africa.

yamanei Smetana, 1984: 130 ( Gabrius ; Type locality: Japan: Gumma Pref.: 5 km E Usui Pass , 900 m).

- DISTRIBUTION: Japan.

zerchei Schillhammer, 1996: 65 ( Gabrius ; Type locality: Russia: Primorskiy Krai Ryazanovka , 14 km SW Slavyanka, 42.48 N, 131.12 E, 50 m).

- DISTRIBUTION: Russia.

Gabronthus

[39 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan,

Oriental, Australian, and Oceanic Regions]

Gabronthus Tottenham, 1955: 180 (species included: maritimus , alluaudanus , pluvialis , thermarum , kisiranus , oribates , tenuissimus , minutissimus , turneri , gombanus , kosanus chyuluensis , fundator , mgogoricus , badalus , meridionalis , styx , inclinans , corrugatus , thermaroides , sulcifrons , pygmaeus , minusculus ). TYPE SPECIES: Gabrius maritimus Motschulsky , fixed by original designation.

— Tottenham, 1955: 181, 190 (keys to African and Oriental species).

— Smetana, 1959: 172 (characters).

— Coiffait, 1963: 7, 12 (characters; notes; thermarum cited as type species).

— Lohse, 1964: 186 (characters; key to central European species).

— Moore, 1965: 97, 99 (characters; notes).

— Shibata, 1973c: 60 (checklist of species of Taiwan).

— Bordoni, 1973j: 707 (list of species from islands near Sicily).

— Coiffait, 1974: 111 (characters; key to species of western Palaearctic region).

— Moore and Legner, 1975: 15 (catalog; North America north of Mexico).

— Pope, 1977: 30 (list of British species).

— Moore and Legner, 1979: 21 (characters; notes).

— Muona, 1979: 15 (list of Scandinavian species).

— Burakowski, Mroczkowski, and Stefańska, 1980: 79 (catalog; Poland).

— Shibata, 1983: 124 (checklist of species of Japan).

— Tóth, 1984: 80 (characters; habitus; key to species of Hungary).

— Frank, 1986a: 370 (checklist of species of Florida).

— Drugmand 1987b (comments on distribution of Belgian species).

— Ádám, 1987: 151 ( Gabronthus ; collecting notes; Hungary).

— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Smetana, 1993b: 48 (list of species of Czech Republic and Slovakia).

— Smetana, 1995e: 478 (characters; key to North American species; notes).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 29 (additional records to Danish list of species).

— Ciceroni and Zanetti, 1995: 29 (list of species of Italy).

— M. Hansen, 1996: 37, 106 (probably belongs within Philonthus ; list of Danish species).

— Downie and Arnett, 1996: 383 (characters).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

— Newton, Thayer, Ashe, and Chandler, 2000: 396 (2 Nearctic species; notes; characters in key). allardi Levasseur, 1964: 291 ( Gabronthus ; Type locality: Luena, Haut Lomami, Katanga).

- DISTRIBUTION: Congo.

alluaudanus Jeannel and Paulian, 1945: 90 ( Philonthus ; subgenus Gabrius ; Type locality: Diego Suarez).

— Tottenham, 1955: 183 ( Gabronthus ; characters).

- DISTRIBUTION: Madagascar.

angustatus Kraatz, 1859 , see: thermarum Aubé, 1850 .

australicus Cameron, 1943a: 343 ( Philonthus ; subgenus Gabrius ; Type locality: Australia: Port Darwin).

— Schillhammer, 1999a: 62 ( Gabronthus ).

- DISTRIBUTION: Australia.

badalus Tottenham, 1955: 186 ( Gabronthus ; Type locality: S. Nigeria).

- DISTRIBUTION: Nigeria.

balthasari Smetana, 1956 , see: limbatus Fauvel, 1900 .

chejuensis Cho, 1996: 28 ( Gabronthus ; Type locality: Korea: Chejudo , Kyoraeri).

- DISTRIBUTION: Korea.

chyuluensis Cameron, 1942b: 329 ( Philonthus ; subgenus Gabrius ; Type locality: Chyulu Hills, altitude 5600 feet).

— Cameron, 1950g: 42 ( Philonthus ; subgenus Gabrius ; Belgian Congo).

— Tottenham, 1955: 189 ( Gabronthus ; characters).

— Tottenham, 1962: 141 ( Gabronthus ; Tanganyika).

- DISTRIBUTION: Kenya, Tanzania, Congo.

corrugatus Tottenham, 1955: 194 ( Gabronthus ; Type locality: Philippines Islands : Luzon, Los Banos).

— Last, 1968: 368 ( Philonthus ; [Note: Misidentified as thermarum according to Last, 1981: 125]; New Guinea).

— Last, 1981: 125 ( Gabronthus ; New Guinea).

- DISTRIBUTION: Philippines, New Guinea.

exilis Kraatz, 1851 , see: thermarum Aubé, 1850 .

flavicollis Motschulsky, 1858: 660 ( Heterothops ; Type locality: Indes orientales).

— Bernhauer and Schubert, 1916: 412 ( Heterothops ; catalog).

— Cameron, 1932: 266 ( Heterothops ; characters; Indes orientales).

— Scheerpeltz, 1933: 1422 ( Heterothops ; catalog).

— Cameron, 1934d: 79 ( Heterothops ; type examined; synonym of Gabrius pulchellus ).

— Smetana, 1988: 319 ( Gabronthus ).

- DISTRIBUTION: “East Indies”, Sri Lanka, Bhutan, Malaysia, Singapore, Indonesia. pulchellus Kraatz, 1859: 92 ( Philonthus ; [preoccupied]; Type locality: India orientalis; Ceylan).

— Bernhauer and Schubert, 1914: 351 ( Philonthus ; catalog).

— Cameron, 1921a: 378, 405 ( Philonthus ; characters; Singapore).

— Cameron, 1932: 136 ( Philonthus ; characters; Ceylon).

— Scheerpeltz, 1933: 1356 ( Philonthus ; catalog).

— Cameron, 1933d: 387 ( Philonthus ; Sumatra).

— Coiffait, 1977b: 206 ( Gabronthus ; Bhutan).

— Smetana, 1988: 319 ( Gabronthus ; synonym of flavicollis ).

foveifrons Cameron, 1932a: 142 ( Philonthus ; subgenus Gabrius ; Type locality: Selangor: Bukit Kutu. Pahang: Raub, Bentong).

— Schillhammer, 1999a: 62 ( Gabronthus ).

fundator Tottenham, 1955: 190 ( Gabronthus ; Type locality: Kenya: Muguga ).

— Tottenham, 1962: 141, 235 ( Gabronthus ; characters; Kenya).

- DISTRIBUTION: Kenya.

fuscolaterus Motschulsky, 1860 , see: thermarum Aubé, 1850 .

gombanus Tottenham, 1955: 188 ( Gabronthus ; Type locality: Abyssinia: Djem-Djem Forest ).

— Tottenham, 1962: 141 ( Gabronthus ; Tanganyika).

- DISTRIBUTION: Ethiopia, Tanzania.

gracilis Coiffait, 1983c: 181 ( Gabronthus ; Type locality: Tahiti: Mont Arue).

- DISTRIBUTION: Society Islands.

inclinans Walker, 1859: 51 ( Xantholinus ; Type locality: Ceylon).

— Fauvel, 1903d: 271 ( Philonthus ; synonym of thermarum ).

— Fauvel, 1905e: 83 ( Philonthus ; synonym of thermarum ).

— Bernhauer and Schubert, 1914: 357 ( Philonthus ; synonym of thermarum ).

— Cameron, 1932: 134 ( Philonthus ; synonym of thermarum ).

— Blackwelder, 1943: 403 ( Philonthus ; synonym of thermarum ).

— Tottenham, 1955: 191 ( Gabronthus ; valid species; characters; angustatus Kraatz may be synonym; Ceylon; Sumatra; Java).

— Scheerpeltz, 1957c: 234 ( Philonthus ; [Note: I assume the author overlooked Tottenham’s elevation of the species]; synonym of thermarum ).

— Smetana, 1958: 217 ( Philonthus ; [Note: I assume the author overlooked Tottenham’s elevation of the species]; synonym of thermarum ).

— Coiffait, 1974: 114 ( Gabronthus ; [Note: I assume the author overlooked Tottenham’s elevation of the species]; synonym of thermarum ).

— Scheerpeltz, 1978: 224 ( Philonthus ; [Note: I assume the author overlooked Tottenham’s elevation of the species]; synonym of thermarum ).

- DISTRIBUTION: Sri Lanka, Indonesia.

kisiranus Tottenham, 1955: 187 ( Gabronthus ; Type locality: Senegal).

— Tottenham, 1961: 205 ( Gabronthus ; Senegal).

- DISTRIBUTION: Senegal.

kosanus Tottenham, 1955: 189 ( Gabronthus ; Type locality: Kenya: Muguga ).

- DISTRIBUTION: Kenya.

levasseuri Jarrige, 1978: 288 ( Gabronthus ; Type locality: Andringitra Centre, plateau Andohariana , 2000–2100 m).

- DISTRIBUTION: Madagascar.

libanicus Saulcy, 1864 , see: maritimus Motschulsky, 1858 .

limbatus Fauvel, 1900d: 230 ( Philonthus ; Type locality: Baical, rivière Sludianka).

— Bernhauer and Schubert, 1914: 344 ( Philonthus ; catalog).

— Scheerpeltz, 1933: 1349 ( Philonthus ; catalog).

— Coiffait, 1974: 116 ( Gabronthus ; characters; Siberia ).

— Schuh, Schillhammer, and Zettel, 1992: 220 ( Gabronthus ; notes; Austria).

— Smetana, 1993b: 48 ( Gabronthus ; Slovakia).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabronthus ; note; Austria).

- DISTRIBUTION: Russia, Slovakia, Austria.

balthasari Smetana, 1956b: 171 , 173 ( Gabrius ; Type locality: Ostslowakei bei Královský Chl’mec).

— Smetana, 1957: 58 ( Gabrius ; characters; Slovakia).

— Smetana, 1959e ( Gabronthus ; characters).

— Smetana, 1958: 257 ( Gabrius ; characters; ecology; Czechoslovakia).

— Lohse, 1964: 187 ( Gabronthus ; characters; central Europe).

— Coiffait, 1974: 115 ( Gabronthus ; cited as balthazari; characters; Slovakia).

— Tóth, 1984: 82 ( Gabronthus ; cited as balthazari; characters).

— Lucht, 1987: 106 ( Gabronthus ; checklist; Czechoslovakia).

— Smetana, 1993b: 48 ( Gabronthus ; synonym of limbatus ).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabronthus ; synonym of limbatus ).

lineifrons Cameron, 1926b: 359 ( Philonthus ; subgenus Gabrius ; Type locality: Simla and Chakrata Districts, 7000 to 8000 feet).

— Cameron, 1932: 135 ( Philonthus ; characters; India).

— Scheerpeltz, 1933: 1349 ( Philonthus ; catalog).

— Schillhammer, 1999a: 62 ( Gabronthus ).

- DISTRIBUTION: India.

maritimus Motschulsky, 1858: 661 ( Gabrius ; Type locality: Alexandrie en Egypte).

— Gemminger and Harold, 1868: 594 ( Philonthus ; synonym of thermarum ).

— Fauvel, 1869: 489 ( Philonthus ; valid species).

— Fauvel, 1874: 245 [= 1874a: 471] ( Philonthus ; race of thermarum ; characters; France; Sicily; Corsica; Algeria; Cyprus; Egypt; Syria).

— Fauvel, 1878: 125 [= 1878a: 45] ( Philonthus ; variety of thermarum ; Algeria; Egypt).

— Fauvel, 1886: 56 [= 1886a: 48] ( Philonthus ; variety of thermarum ; notes; Morocco; Egypt; Algeria).

— Ganglbauer, 1895: 460 ( Philonthus ; subgenus Gabrius ; characters; middle Europe).

— Fauvel, 1902b: 111 ( Philonthus ; Morocco; Algeria; Egypt).

— Sainte-Claire Deville, 1906: 98 ( Philonthus ; Corsica).

— Bernhauer and Schubert, 1914: 345 ( Philonthus ; catalog).

— Cameron, 1921a: 378, 405 ( Philonthus ; characters; Singapore).

— Portevin, 1929: 366 ( Philonthus ; subspecies of thermarum ; characters; France).

— Gridelli, 1930: 74 ( Philonthus ; Libya).

— Bernhauer, 1931d: 587 ( Philonthus ; Abyssinia).

— Scheerpeltz, 1933: 1350 ( Philonthus ; catalog).

— C. Koch, 1941: 63 ( Philonthus ; Morocco).

— Bernhauer, 1942c: 361 ( Philonthus ; Cameroon).

— Jarrige, 1951: 134 ( Philonthus ; Annam; Indochina).

— Smetana, 1953: 119 ( Philonthus ; Turkey).

— Tottenham, 1955: 185 ( Gabronthus ; characters; many species are misidentified as maritimus ). — Smetana, 1959: 173 ( Gabronthus ; characters).

— Scheerpeltz, 1963b: 429 ( Philonthus ; subgenus Gabronthus ; Egypt).

— Smetana, 1967i: 555 ( Gabronthus ; Turkey) .

— Coiffait, 1973a: 274 ( Philonthus ; Morocco).

— Coiffait, 1974: 113 ( Gabronthus ; characters; Mediterranean region).

— Watanabe and Shibata, 1976: 332 ( Philonthus ; subgenus Gabronthus ; Malaysia).

— Scheerpeltz, 1978: 224 ( Philonthus ; subgenus Gabronthus ; Timor).

— Smetana, 1978: 116 ( Gabronthus ; Israel).

— Coiffait, 1981b: 236 ( Gabronthus ; Saudi Arabia).

— Coiffait, 1986: 393 ( Gabronthus ; Saudi Arabia).

— Lecoq, 1990: 196 ( Gabronthus ; Mascarene Islands).

— Ito, 1992a: 64 ( Gabronthus ; Yakushima Island).

— Shibata, 1993a: 318 ( Gabronthus ; Taiwan).

— Ciceroni and Zanetti, 1995: 29 ( Gabronthus ; Italy).

— Gusarov, 1997: 287 ( Gabronthus ; Saudi Arabia).

— Assing, 1998e: 140 ( Gabronthus ; Canary Islands).

— Assing, 2000b: 115 ( Gabronthus ; collecting notes; Canary Islands).

- DISTRIBUTION: France, Italy, Cyprus, Turkey, Cameroon, Canary Islands, Morocco, Algeria, Libya, Egypt, Ethiopia, Saudi Arabia, Israel, Lebanon, Syria, Mascarene Islands, Malaysia, Vietnam, Singapore, Indonesia, Taiwan, Japan.

libanicus Saulcy, 1864: 639 ( Philonthus ; Type locality: Beïroût).

— Baudi, 1870: 386 ( Philonthus ; characters; Cyprus; Lebanon).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of maritimus ).

— Bernhauer and Schubert, 1914: 346 ( Philonthus ; synonym of maritimus ).