Philisca viernes, Soto & Ramírez, 2012

Soto, Eduardo M. & Ramírez, Martín J., 2012, Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443, Zootaxa 3443 (1), pp. 1-65 : 46-50

publication ID	https://doi.org/ 10.11646/zootaxa.3443.1.1
persistent identifier	https://treatment.plazi.org/id/F31587CF-A61B-7954-FF6E-1CD20259FC50
treatment provided by	Felipe
scientific name	Philisca viernes
status	sp. nov.

Treatment

Philisca viernes View in CoL new species

Figs 1f View FIGURE 1 , 28–30 View FIGURE 28 View FIGURE 29 View FIGURE 30 , 31b View FIGURE 31 , 33c, d View FIGURE 33 , 35e View FIGURE 35

Type material: Holotype male and female paratype from Chile, Región V (Valparaíso), Valparaíso Prov., Archip. Juan Fernández, Is. Robinson Crusoe, sector Salsipuedes, S33.63087° W78.84386° (GPS, área +- 100m), elev. 482m (barómetro), 21.II.2011, bosque de canelo, maqui y luma, M. Ramírez, E. Soto, J. Pizarro ( MHNS; temporary preparations: male EMS-00152, female EMS-00151); same locality, 1 female paratype, Anson valley, Plazoleta de El Yunque , 1–28.IV.1962, B. Malkin ( AMNH: female EMS-00052). GoogleMaps

Etymology: The specific name is a noun in apposition, taken from the name of the indigenous friend of Robinson Crusoe in the novel of Daniel Defoe.

Diagnosis: It resembles other species of the ornata group by the modified male chelicerae and the epigyne with a smooth, heart shaped median field close to the epigastric furrow ( Figs 35e View FIGURE 35 , 33c, d View FIGURE 33 ). It can be distinguished by having few or no spines on tibiae I and II (up to two ventral spines).

Description: Female (paratype MHNS EMS-00151): Total length 8.38. Prosoma length 4.12, width 2.93, wider between legs II–III. Tibia/metatarsus length: I, 2.13/1.67; II, 2.00/1.53; III, 1.33/2.13; IV, 2.13/3.00. Chelicerae with three teeth (middle tooth larger) on promargin and four on retromargin (apical tooth smaller). Sternum length 1.93, width 1.53. Spines: leg I, femur d 1–1–1, p 2ap. II, femur d 1–1–1, p 1ap; metatarsus v 2bas. III, femur d 1–1–1, p d1ap or 0–d1–d1, r d1ap; patella r d1; tibia v p1–2–2, p and r d1–1; metatarsus v 2–p1–2 or 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p 1–1, r d1–1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Opisthosoma length 4.26, epigastrium–spiracle 2.40, spiracle– spinnerets 0.47. Color in ethanol: Prosoma and chelicerae reddish brown. Sternum brown. Legs yellowish brown. Opistosoma light yellowish brown, with dorsal and ventral brown pattern ( Figs 1f View FIGURE 1 , 28 View FIGURE 28 ). Epigyne small, very sclerotized and weakly translucent ( Fig. 33c, d View FIGURE 33 ). MF smooth, heart-shaped, close to the epigastric furrow. Anterior pouch wide, with lumen shorter than wide. AB placed anteriorly from the spermathecae, with long ducts. Spermathecae far apart ( Fig. 33d View FIGURE 33 ).

Male (holotype): Total length 7.45. Prosoma length 3.60, width 2.66. Length of tibia/metatarsus: I, 2.60/2.20;

II, 2.47/2.33; III, 1.87/2.40; IV, 2.60/3.33. Chelicerae and endites with clear modifications ( Figs 29 View FIGURE 29 ; 35e View FIGURE 35 ). Chelicerae with three teeth (middle tooth larger) on promargin, three or four on retromargin. Sternum length 1.93, width 1.47. Spines: leg I, femur d 1–1–1, p d2ap; metatarsus v 2bas–p1. II, femur d 1–1–1, p d2ap or d1ap, r d1ap; metatarsus v 2bas–p1 or 2–2–0. III, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–p1–2, p and r d1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p d1– 1, r d1–d1–0–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Opisthosoma length 3.73, spiracle–epigastrium 1.87, spiracle–spinnerets 0.47. Color in ethanol ( Fig. 29 View FIGURE 29 ): Prosoma and chelicerae reddish brown. Sternum brown. Legs yellowish brown. Opistosoma light yellowish brown, with dorsal and ventral brown pattern. Palp ( Fig. 30 View FIGURE 30 ): Tibia width/length 0.36. Embolus long, thin and cylindrical. MA long and flattened, with a few thin small branches ( Fig. 30h View FIGURE 30 ). PMA long and wide, triangular, pPMA not simple. Conductor large and complex, with vestigial canal, ending in longitudinally ridged peak, on rounded projection, directed backwards ( Fig. 30 View FIGURE 30 ).

All adults and immatures have a group of thick setae on the base of ALS ( Fig. 31b View FIGURE 31 ).

Variability: Spines in females: II, femur r 1ap; metatarsus v d1bas. III, tibia v p1–1–0; metatarsus r d1–0–1. IV, tibia, v p1–p1–0.

Distribution: Known only from the Juan Fernández Archipelago, Isla Robinson Crusoe, Chile.

Other material examined: CHILE: Región V (Valparaíso): Valparaíso Prov., Archip. Juan Fernández , Is. Robinson Crusoe, sendero, luego de Mirador Selkirk , ladera W, S33.63790° W78.85302° (GPS, trayecto +- 110m), elev. 460–470m (barómetro), 20.II.2011, bosque de luma con helechos arborescentes, M. Ramírez, E. Soto, J. Pizarro GoogleMaps , 1 male ( MACN-Ar 27965 ) , 1 immature ( MACN-Ar 28085 ) ; Sector Salsipuedes (same locality as holotype), S33.63087° W78.84386° (GPS, área +- 100m), elev. 482m (barómetro), 21.II.2011, bosque de canelo, maqui y luma, M. Ramírez, E. Soto, J. Pizarro GoogleMaps , 1 female ( MACN-Ar 28087 ) , 1 male ( MACN-Ar 27966 ) , 1 male ( UC: MJR-loc-13/2) , 1 immature ( MACN-Ar 27967 ) , 1 immature ( MACN-Ar 28086 ) ; Plazoleta de El Yunque , 1–28.IV.1962, B. Malkin , 1 female ( AMNH) .