Peltonotus, Burmeister

Jameson, Mary Liz & Wada, Kaoru, 2004, Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia, Zootaxa 502 (1), pp. 1-66 : 9-11

Key to Male Peltonotus Species

(Males of P.kyojinus , P. nethis , P. pruinosus , and P. suehirogarus are not known)

1. Apical half of mentum acute, triangulate ( Figs. 19, 28 ) ............................................... 2

1'. Apical half of mentum rounded ( Figs. 20–23, 25–27 ) or quadrate ( Fig. 24 )................ 3

2. Smaller foretarsal claw deeply arcuate ( Fig. 33 ); genitalia as in Fig. 39 ........................ ......................................................................... P. deltamentum Jameson & Wada , n. sp.

2'. Smaller foretarsal claw simply arched; genitalia as in Fig. 49 ........................................ ................................................................................... P. sisyrus Jameson & Wada , n. sp.

3. Apex of labrum bilobed ( Figs. 16–17 ).......................................................................... 4

3'. Apex of labrum broadly emarginate ( Figs. 15, 18 ) to deeply, narrowly emarginate ( Fig. 14 ) ................................................................................................................................. 5

4. Foretibia tridentate, basal tooth well­developed; genitalia as in Fig. 45 ......................... .............................................................................................................. P. nasutus Arrow

4'. Foretibia bidentate, basal tooth poorly developed; genitalia as in Fig. 44 ...................... ......................................................................................................... P. morio Burmeister

5. Labrum with apex deeply, narrowly emarginate (e.g., Fig. 14 ) .................................... 6

5'. Labrum with apex broadly emarginate ( Figs. 15, 18 ) ................................................... 9

6. Mala of maxilla with setae thick and strongly flattened; occurring in Borneo, Malaysia, and Sumatra; genitalia as in Figs. 38 or 41............................................................. 7

6'. Mala of maxilla with setae not thick and strongly flattened; occurring in South Vietnam; genitalia as in Fig. 42 ........................................................... P. karubei Muramoto

7. Punctures of frons lacking setae; genitalia as in Fig. 38 ..... P. brunnipennis Benderitter

7'. Punctures of frons with short, velutinous setae and moderately long setae; genitalia not as in Fig. 41 ................................................................................................................... 8

8. Fifth foretarsal segment and claw gracile; maxillary stipes with setae curly at apex (e.g., Fig. 30 ); Sarawak.................................... P. gracilipodus Jameson & Wada , n. sp.

8'. Fifth foretarsal segment and claw greatly thickened, robust; maxillary stipes with setae straight, not curly at apex; Malaysia (Cameron Highlands)............................................ .......................................................................... P. podocrassus Jameson & Wada , n. sp.

9. Labial palpomere 2 greatly enlarged and dorsoventrally flattened, 2–3 times wider than palpomere 1 ( Fig. 22 ) .......................................................................................... 10

9'. Labial palpomere 2 not greatly enlarged and flattened, less than 1.5 times wider than palpomere 1 ( Fig. 27 ) .................................................................................................. 11

10. Maxillary stipes with setae curly at apex ( Fig. 30 ); genitalia as in Fig. 43 ..................... ........................................................................................................ P. malayensis Arrow

10'. Maxillary stipes with setae straight, not curly at apex; genitalia as in Fig. 47 ............... ................................................................................. P. silvanus Jameson & Wada , n. sp.

11. Foretibia tridentate; genitalia as in Fig. 40 ............... P. fujiokai Jameson & Wada , n. sp.

11'. Foretibia bidentate ( Fig. 31 ); genitalia not as in Fig. 40 ............................................. 12

12. Elytra reddish with castaneous vittae ( Figs. 12–13 ); genitalia as in Fig. 50 ................... .............................................................................................................. P. vittatus Arrow

12'. Elytra lacking vittae, entirely castaneous or black; genitalia not as in Fig. 50 ........... 13

13. Pronotal basal bead lacking; length less than 12.0 mm; genitalia as in Fig. 46 .............. ................................................................................... P. rubripennis Miyake & Yamaya

13'. Pronotal basal bead extending to beyond basal angle; length greater than 17.0 mm; genitalia not as in Fig. 46 ............................................................................................ 14

14. Fifth foretarsomere with well­developed apical protrusion anteriomedially ( Fig. 36 ), lacking weak medial protrusion; occurring in region surrounding Mt. Bawang , Kalimantan (Fig. 67) ........................................... P. adelphosimilis Jameson & Wada , n. sp.

14'. Fifth foretarsomere lacking apical protrusion; weak protrusion at middle ( Fig. 35 ); occurring in Sabah (Fig. 67) .................................................................. P. similis Arrow

Figures

FIGURES 19–28. Mentum, dorsal view of: 19) P. deltamentum, 20) P. gracilipodus, 21) P. kyojinus, 22) P. malayensis, 23) P. podocrassus, 24) P. nasutus, 25) P. nethis, 26) P. pruinosus, 27) P. similis, 28) P. sisyrus.

FIGURES 31–36. Detail of right forelegs of: 31) P. malayensis (showing bidentate foretibia, dorsal view), 32) P.rubripennis (showing tridentate foretibia with basal tooth very weakly developed, dorsal view), 33) P. deltamentum (showing strongly arcuate smaller claw, ventral view), 34) P. nasutus (showing form of foretarsomeres 1–5), 35) P.similis (showing form of foretarsomeres 1–5 and median protuberance of foretarsomere 5 [arrow]), 36) P. adelphosimilis (showing form of foretarsomeres 1–5 and apical protuberance of foretarsomere 5 [arrow]).

FIGURES 37–39. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 37) P. adelphosimilis, 38) P. brunnipennis, 39) P. deltomentum.

FIGURES 49–50. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 49) P. sisyrus, 50) P. vittatus.

FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally).

FIGURES 43–45. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 43) P. malayensis, 44) P.morio, 45) P. nasutus.

FIGURES 40–42. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 40) P.fujiokai, 41) P. gracilipodus and P. podocrassus, 42) P.karubei.

FIGURES 29–30. Maxilla (dorsal view) showing articulated maxillary tooth, mala with lamellate setal brush (black arrow), and form in: 29) P. deltamentum and 30) P. malayensis.

FIGURES 46–48. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 46) P.rubripennis, 47) P. silvanus, 48) P. similis.

FIGURES 2–13. Dorsal habitus of: 2–4) P. fujiokai (males), 5) P. fujiokai (female), 6) P. malayensis (male), 7) P. malayensis (female), 8) P. morio (male), 9) P. morio (female), 10) P. nasutus (male), 11) P.nasutus (female), 12) P. vittatus (male), 13) P. vittatus (female).