Pedethma malandensis Weise

Lingafelter, Steven W. & Konstantinov, Alexander S., 2000, Revision Of Pedethma Weise (Coleoptera: Chrysomelidae), The Coleopterists Bulletin 54 (4), pp. 413-458 : 443-455

Pedethma malandensis Weise

( Figs. 16a , 24a , 25a)

Description. Measurements listed in Table 1. Basal antennomeres pale, apical 3–4 antennomeres darkened. Vertex with two setiferous pores on each side between upper portion of eye forming straight line, but widely separated at middle of vertex. Supracallinal, midfrontal, and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. Several large pores forming two rows along margin of eye between antennal callus and orbit. Orbit extremely narrow, 0.42 times width of antennal callus. Antennal socket situated very close to eye margin. Upper part of frontal ridge very narrow between antennal calli, no more than one­fourth width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, narrower than latter. No ridges between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye ( Fig. 16a ) very large and occupying over two­thirds of head as viewed from lateral perspective in males. Eye 1.87 times higher than wide. Pronotum ( Fig. 16a ) brownish, darker at anterior two­thirds than region posterior of transverse impression. Posterior pronotal depression straight. Elytra ( Fig. 16a ) unicolorous, pale to moderately dark brown. Abdominal venter and metasternum darker than surrounding region. Legs pale yellow­brown. Aedeagus ( Figs. 24a , 25a) symmetrical, not strongly tapering. Apex of aedeagus truncate with slight apical, medial impression and two lateral apical arms. Dorsal transverse ridges present. Aedeagus weakly curved as viewed from lateral perspective. Spermatheca unavailable for description. Tignum of paralectotype with basal sclerotization apparently unique for the genus.

Diagnostic and Other Comments. This species was described by Weise (1923:120). It is distinctive with the very large eyes, narrow frons and frontal ridge, and anteriorly narrowed antennal calli. The truncate aedeagal apex with apico­lateral arms and unique basal sclerotization of the tignum is characteristic. This species differs from P. kurandensis by its amber body color, pale to yellow legs, reduced number of pores above the eye, and larger pronotal punctures.

Distribution. Rainforests of northeast Queensland (Map 1, locality #4).

Material. Lectotype (designated herein): (Male: labels numbered): 1) Malanda; 2) Queensl., Mjoberg; 3) (no data); 4) loan 1132/94; 5) Lectotype Pedethma malandensis Weise ( NHRS) .

Paralectotypes. Labels 1–2 as in lectotype; 3) type; 4) (no data); 5) Typus; 6) Pedethma malandensis m. 7) loan 1133/94; 8) Paralectotype Pedethma malandensis Weise ( NHRS, 1 specimen); Labels 1–2 as in lectotype; 3); 4 ) Paratypus; 5) Pedethma malandensis m. 6) loan 1131/94; 7) Paralectotype Pedethma malandensis Weise ( NHRS, 1 specimen) .

Other Material. ‘‘ NEQ: 17°23'S 145°46'E Bartle Frere, W. Base, 7 Mar­ 15 May 1995, Monteith and Hasenpusch, Pitfall traps, 700 m’’ ( QMBA, 1 specimen); ‘‘NE QLD. Emerald Ck, Lamb Range, 11 Oct 1982, 950 m, Monteith, Yeates and Thompson, Pyrethrum knockdown, RF’’ ( NMNH, 1 specimen) .

Pedethma nigra Lingafelter and Konstantinov , new species ( Figs. 15f , 22k , 28e , 29c )

Description. Measurements listed in Table 1. Antennae over two­thirds length of body in males, slightly more than one­half length of body in females. Basal 4–5 antennomeres pale, apical antennomeres dark brown to piceous. Antennal calli pale yellow. Vertex with two setiferous pores on each side between upper portion of eye, forming straight line. Supracallinal sulcus poorly developed. Midfrontal and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores between antennal callus and orbit. Orbit extremely narrow, one­half width of antennal callus. Antennal socket very close to eye margin. Upper part of frontal ridge between antennal calli one­fifth width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, narrower than latter. No ridges between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye twice as high as wide, slightly higher than distance between eyes (above antennal sockets). Pronotal shape with moderate antemedial expansion and posterior constriction. Posterior pronotal depression sinuate with slight posterior dip at middle and sides. Pronotum pale yellow. Elytra ( Fig. 15f ) dark reddish­brown, appearing black. Abdominal venter and metasternum piceous. Aedeagus ( Fig. 22k ) long, slightly expanded apically, but not as strongly as in P. maculata ; without abrupt preapical constriction as in P. maculata . Aedeagus with very slight apical, medial impression; no other ridges or impressions present. Aedeagus moderately curved as viewed from lateral perspective. Spermatheca ( Figs. 28e , 29c ) abruptly curved at each end, slightly sinuate along body. Proximal and distal ends widely separated by about one­half of overall length of spermatheca.

Diagnostic and Other Comments. The very pale yellow head, pronotum, and legs, small size, dark reddish­brown elytra (appearing black), long aedeagus without any distinct impressions or anteapical constriction, and sinuate spermatheca with widely separated proximal and distal ends distinguish this species from other Pedethma species.

Distribution. Rainforests of northeast Queensland (Map 1, locality #4).

Etymology. The specific epithet refers to the dark elytra.

Material. Holotype ( Male ): ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ,

Summit TV Stn., 1,560 m, 17 Oct.–Nov.5, 1981, EARTHWATCH/ QLD. MU­ SEUM, Malaise trap, rainforest’’ ( ANIC).

Paratypes. ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/ QLD. MUSEUM , Malaise trap, rainforest’’ ( ANIC, 2 specimens); ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTH­ WATCH / QLD. MUSEUM, Beating, rainforest’’ ( ANIC, 1 specimen); ‘‘[AUS­ TRALIA]: Bellenden Ker Range, NQ, Cable Tower , 3, 1,054 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/ QLD. MUSEUM , Beating, rainforest’’ ( ANIC, 3 specimens; NMNH, 3 specimens); ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/ QLD. MUSEUM’’ ( ANIC, 1 specimen); ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/ QLD. MUSEUM , Pyrethrum knockdown’’ ( ANIC, 2 specimens; NMNH, 1 specimen); ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/ QLD. MUSEUM , Pitfall trap, rainforest’’ ( ANIC, 1 specimen; NMNH, 1 specimen); ‘‘[ AUSTRALIA]: Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, Oct. 25–31, 1981, EARTH­ WATCH / QLD. MUSEUM, Q. M. BERLESATE No. 372, 17.16S 145.51E, Rainforest, Sieved litter, A.N.I.C. COLEOPTERA Voucher No. 83­1094,’’ ( ANIC, 1 specimen) .

Other Material. ‘‘ Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/QLD. MUSEUM , Beating, Rainforest’ ’ ( QMBA, 2 specimens); ‘‘ Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, Nov. 1–7, 1981, EARTHWATCH/QLD. MUSEUM ; Yellow pan trap in rainforest’’ ( QMBA, 1 specimen); ‘‘ Bellenden Ker Range, NQ, Summit TV Stn., 1,560 m, 17 Oct. – Nov. 5, 1981, EARTHWATCH/QLD. MUSEUM , Pyrethrum knockdown; A.N.I.C., COLEOPTERA Voucher No. 83­1094’’ ( QMBA, 1 specimen); ‘‘NE.Q: 17°14'S × 145°48'E, Massey Ra., 6 km NW of Bellenden Ker , 1,150 m, 11 Oct 1991. PYRETHRUM, Monteith and Janetzki’ ’ ( QMBA, 1 specimen) .

Pedethma pinnipenis Lingafelter and Konstantinov , new species ( Figs. 16e , 23g , 27c , 31m )

Description. Measurements listed in Table 1. Antennae about two­thirds length of body in males, slightly more than one­half length of body in females. Most antennomeres pale, becomming darker brown at apex of antenna. Antennal calli pale yellow to reddishbrown, of same color as head. Vertex with two to three setiferous pores on each side between upper portion of eye, forming straight line. Supracallinal sulcus well developed. Midfrontal and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending between antennal socket and frontal ridge. A row of large pores between antennal callus and orbit. Orbit extremely narrow, less than one­half width of antennal callus. Antennal socket very close to eye margin. Upper part of frontal ridge between antennal calli less than one­fifth width of transverse diameter of antennal socket. Lower part of frontal ridge forming obtuse angle with upper part, narrower than latter. No ridges between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye 1.9 as high as wide, slightly higher than distance between eyes (above antennal sockets). Pronotal shape with moderate antemedial expansion and posterior constriction. Posterior pronotal depression nearly straight with slight posterior dip at middle and sides. Pronotum reddish­brown, darker at sides. Elytra ( Fig. 16e ) variable, pale yellow­brown with darker epipleura and periscutellar region on holotype and some paratypes to uniformly pale yellow on other paratypes. Abdominal venter and metasternum yellow to reddish­brown. Aedeagus ( Figs. 23g , 27c ) long, with strong arrowhead apex, strongly constricted anteapically. Aedeagus with transverse dorsal ridges. Aedeagus moderately curved as viewed from lateral perspective. Spermatheca ( Fig. 31m ) moderately curved at each end, body more thickened at base; proximal and distal ends moderately separated by about one­third to one­half of overall length of spermatheca.

Diagnostic and Other Comments. The very distinctive arrowhead shaped aedeagus with very strong anteapical constriction readily identifies this species.

Distribution. Rainforests of northeast Queensland (Map 1, locality #2).

Etymology. The specific epithet refers to the abruptly arrow­headed aedeagus.

Material. Holotype (Male): ‘‘ Hugh Nelson Ra , 2.5 km S of Crater N.P., N.Qld., 5 Dec 1988. 1,100 m. Monteith and Thompson , Pyrethrum / Logs and Trees’ ’ ( QMBA, QMT 62962).

Paratypes. ‘‘ Hugh Nelson Ra, 2.5 km S of Crater N.P., N.Qld., 5 Dec 1988. 1100 m. Monteith and Thompson, Pyrethrum / Logs and Trees’ ’ ( QMBA, 1 specimen); ‘‘ Sluice Ck , 9 km WSW Millaa Millaa, NEQ, 5–14 Dec 1988. 1,150 m, Monteith and Thompson , Flight Intercept Trap’ ’ ( QMBA, 2 specimens); ‘‘NE QLD. 1.5 km W of Cape Tribulation (Site 3), 23 Sept.–7 Oct. 1982, 150 m. Monteith, Yeates and Thompson, Baited flight trap, RF’’ ( QMBA, 1 specimen); ‘‘NE QLD. 1.5 km W of Cape Tribulation ( Site 3), 25 Sept 1982, 150 m. Monteith, Yeates and Thompson; QM Berlesate No. 425, 16.05S 145.28E, Rainforest , 150 m, Stick brushing’’ ( QMBA, 1 specimen); ‘‘NE QLD. 1.5 km W of Cape Tribulation (Site 3), 23 Sept.–7 Oct. 1982, 150 m. Monteith, Yeates and Thompson, Rainforest pitfall traps’’ ( QMBA, 1 specimen); ‘‘NE QLD. 1.5 km W of Cape Tribulation ( Site 10), 28 Sept. 1982, 780 m. Monteith, Yeates and Thompson, Pyrethrum knockdown, RF’’ ( QMBA, 1 specimen); ‘‘ North Summit , Bakers Blue Mt., 17 km W Mt Molloy, NQ, 1–18 Jan 1990, 1,000 m, anzses, Flt. Intercept, Site 2’’ ( QMBA, 1 specimens; NMNH, 1 specimen) .

Pedethma pubescens Lingafelter and Konstantinov , new species ( Figs. 14f , 21 , 22e , 26b, 28i , 29e , 33d )

Description. Measurements listed in Table 1. Antennae nearly length of body in male, slightly shorter in females. Basal antennomeres pale, apical antennomeres dark reddishbrown to piceous. Antennal calli similar in coloration to surrounding portions of head. Vertex with two setiferous pores on each side between upper portion of eye, forming straight line. Midfrontal and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal sulcus poorly developed. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores separate callus and orbit. Orbit extremely narrow, 0.54 times width of antennal callus. Antennal socket situated very close to eye margin. Upper part of frontal ridge between antennal calli 0.48 times width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, narrower than latter. No narrow ridges situated between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye size of males extremely large and protuberant, occupying about two­thirds of head as viewed from lateral perspective. Of females, eyes smaller, but still protuberant. Pronotum ( Figs. 14f , 21 ) densely pubescent with long hairs, broad, with moderate antemedial expansion and posterior constriction. Posterior pronotal depression nearly horizontal with only slight posterior tapering at ends. Pronotum pale yellowish. Elytra ( Figs. 14f , 21 ) densely pubescent with long hairs, dark reddish­brown, appearing black. Abdominal venter dark brown; metasternum reddish­brown. Aedeagus ( Figs. 22e , 26b) long, tapering continuously to narrow point at apex (as in P. kurandensis ) with transverse dorsal ridges and minute ventral striae present at middle third. Aedeagus strongly curved with very narrow apical fifth as viewed from lateral perspective. Spermatheca ( Figs. 28i , 29e ) moderately, symmetrically curved at distal and proximal ends (as in P. howdeni and P. suturalis ) with separation approximately one­third length of spermatheca.

Diagnostic and Other Comments. Although this species is extremely similar to P. kurandensis in the shape of the aedeagus, it is easily recognized by its abundant and uniformly distributed long setae on the elytra and pronotum.

Distribution. Rainforests of northeast Queensland (Map 1, locality #2).

Etymology. The specific epithet refers to the hairy elytra and pronotum.

Material. Holotype ( Male ): ‘‘[ AUSTRALIA]: Bamboo Ck, near Miallo, N. of Mossman, N. Qld., 25 Apr. 1967, D. H. Colless’ ’ ( ANIC).

Paratypes. ‘‘[ AUSTRALIA]: Bamboo Ck , near Miallo, N. of Mossman, N. Qld., 25 Apr. 1967, D. H. Colless’ ’ ( ANIC, 3 specimens; NMNH, 1 specimen) .

Pedethma seymourensis Lingafelter and Konstantinov , new species ( Figs. 16g , 27f , 31o )

Description. Measurements listed in Table 1. Antennae about two­thirds length of body in both sexes. In holotype and allotype, basal 4–5 antennomeres pale, apical antennomeres dark brown to piceous; in other material, most antennomeres are uniformly reddish­brown. Antennal calli pale yellow but similar in color to surrounding head. Vertex with two or three setiferous pores of unequal size on each side between upper portion of eye, not forming very straight line. Supracallinal sulcus poorly developed. Midfrontal and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores between antennal callus and orbit. Orbit narrow, nearly as wide as antennal callus. Antennal socket very close to eye margin. Upper part of frontal ridge between antennal calli one­fifth width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, narrower than latter. Area lateral of lower part of frontal ridge slightly swollen. No ridges between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye 2.2 times as high as wide, slightly higher than distance between eyes (above antennal sockets). Pronotal shape with moderate antemedial expansion and posterior constriction; more pronounced in male than female. Posterior pronotal depression straight to slightly recurved. Pronotum pale yellow. Elytra ( Fig. 16g ) pale with piceous epipleura in holotype and allotype; dark periscutellar region in other specimens. Abdominal venter piceous in holotype and allotype; reddish­brown in others; metasternum reddish­brown. Aedeagus ( Fig. 27f ) long, moderately tapered apically, but not as strongly as in P. pubescens or P. kurandensis . Aedeagus with dorsal transverse ridges and rounded apex. Aedeagus moderately curved as viewed from lateral perspective. Spermatheca ( Figs. 31o ) nondistinct, curved at each end but more strongly at apex of body. Proximal and distal ends moderately separated by about one­third of overall length of spermatheca.

Diagnostic and Other Comments. The aedeagus, lacking an arrowhead apex, and with the dorsal, but not ventral transverse ridges is most diagnostic for this species.

Distribution. Rainforests of northeastern Queensland (Map 1, localities #4, 5).

Etymology. The specific epithet refers to the type locality.

Material. Holotype ( Male , on same point with allotype): ‘‘[ AUSTRALIA]: NE.Q: 17°28'S 146°02'E, Polly Ck, Seymour Ra. 12 Sept–22 Oct 1991. 80 m, Monteith and Janetzki, Pitfall and Intercept traps’’ ( QMBA, QMT 62970).

Paratype (Allotype). ‘‘NE.Q: 17°28'S 146°02'E, Polly Ck, Seymour Ra. 12 Sept–22 Oct 1991. 80 m, Monteith and Janetzki, Pitfall and Intercept traps’’ ( QMBA, 1 female specimen, on same point with holotype) .

Other Material. ‘‘NEQ: 17°26'S 145°42'E, Hughes Road, Topaz. 6 Dec 1993 – 25 Feb 1994. Monteith , Cook, Janetzki, RF Pitfalls, 650 m. ’’ ( QMBA, 1 specimen); ‘‘NEQ: 17°35'S 145°35'E, Maalan SF on Hwy. 25 Nov 1994 – 10 Jan 1995. Monteith and Hasenpusch, Flt. intercept trap, 850 m. ’’ ( QMBA, 1 specimen) .

Pedethma sinuatipenis Lingafelter and Konstantinov , new species ( Figs. 16b , 23a–b , 27a )

Description. Measurements listed in Table 1. Male with antennae slightly shorter than body (females unknown). Antennae reddish brown to piceous at apex. Antennal calli not noticeably paler than surrounding region of head. Vertex with three setiferous pores, of differing size, on each side between upper portion of eye, not forming straight line. Supracallinal sulcus poorly developed. Midfrontal and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores between antennal callus and orbit. Orbit extremely narrow, one­half width of antennal callus. Antennal socket very close to eye margin. Upper part of frontal ridge between antennal calli one­fifth width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, narrower than latter. No ridges between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye twice as high as wide, slightly higher than distance between eyes (above antennal sockets). Pronotal shape nearly quadrate with very weak antemedial expansion and posterior constriction. Posterior pronotal depression nearly straight with slight posterior dip at sides. Pronotum pale reddish­brown. Elytra ( Fig. 16b ) diffusely darkened at epipleura and base. Abdominal venter and metasternum reddish­brown. Aedeagus ( Figs. 23a–b , 27a ) long, slightly tapering apically. Aedeagus without dorsal or ventral transverse ridges present. Aedeagus strongly sinuate as viewed from lateral perspective. Spermatheca unknown.

Diagnostic and Other Comments. The relatively long and very strongly sinuate aedeagus is most characteristic for this species. The nearly quadrate pronotum with weak antemedial expansion is also distinctive.

Distribution. Rainforests of northeastern Queensland (Map 1, locality #4).

Etymology. The specific epithet refers to the serpentine curvature of the aedeagus when viewed from lateral perspective.

Material. Holotype (Male): ‘‘ Mt Bartle Frere , N.Qld. Sth Peak Summit, 1,620 m, 6–8 Nov., 1981. EARTHWATCH/ QLD. MUSEUM, Pyrethrum knockdown; A.N.I.C., Voucher No. 83–1095’’ ( QMBA, QMT 62972).

Pedethma suturalis Weise ( Figs. 14a , 25c, 28j , 29f , 32a , 33c )

Description. Measurements listed in Table 1. Dimorphic antennal lengths: males have antennae approximately the length of the body with the apical antennomere widest just distal to middle; females have antennae less than two­thirds length of body and widest portion of apical antennomere is at distal one­third or distad to this point. Basal antennomeres reddish brown, apical antennomeres darker brown to piceous. Antennal calli not noticeably paler than surrounding region of head. Vertex with three setiferous pores on each side between upper portion of eye forming straight line. Supracallinal, midfrontal, and supraantennal sulci well developed. Supraorbital sulcus absent. Supracallinal and midfrontal sulci straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores separate antennal callus and orbit. Orbit narrow, 0.58 times width of antennal callus. Antennal socket situated close to eye margin. Upper part of frontal ridge between antennal calli 0.29 times width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, slightly narrower than latter. No ridges situated between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye 1.94 times higher than wide. Pronotal margin strongly produced anterolaterally, with bulge extending at least as far as anterior corners. Posterior pronotal depression nearly horizontal with only slight posterior dip at middle. Posterior pronotal margin moderately constricted forming a very sinuate margin. Pronotum uniformly reddish brown. Elytra ( Fig. 14a ) with distinct periscutellar patch that tapers along suture. Elytral epipleura black to brown. Color of venter of abdomen piceous. Color of metasternum piceous. Aedeagus ( Fig. 25c) slightly tapering apically. Apex of aedeagus truncate, not constricted (as in P. malandensis ). A slight longitudinal impression extending approximately one­third length of aedeagus from apex. Aedeagus thick and moderately curved from lateral perspective. Spermatheca ( Figs. 28j , 29f , 32a ) broadly curved distally as in P. howdeni , with a wide separation between distal and proximal ends. Surface sculpture of spermatheca distinctly imbricate ( Fig. 32a ).

Diagnostic and Other Comments. This species was described by Weise (1923:120–121). It is a distinctively maculate species with the periscutellar maculation extending along suture to the elytral apex, and possessing widely produced pronotal margins. Less maculate forms resemble P. howdeni , but the aedeagi differ between the two significantly. The spermathecae of both species are similar in shape, but the surface sculpturing varies: In P. suturalis it is imbricate, in P. howdeni it is fissate.

Distribution. Rainforests of northeast Queensland (Map 1, locality #4).

Material. Lectotype (designated herein): (Male: labels numbered): 1) Herberton; 2) Queensl., Mjoberg; 3) (no data); 4) type; 5) Typus; 6) Pedethma suturalis m.; 7) Lectotype Pedethma suturalis Weise des. A. Konstantinov, 1995 ( NHRS) .

Paralectotype. Labels 1–3 as in lectotype; 4) loan 1135/94; 5) Paralectotype Pedethma suturalis Weise des. A. Konstantinov, 1995 ( NHRS, 1 specimen) .

Other Material. ‘‘ AUSTRALIA, Atherton, Qld., 8.ll.1975, H. and A. Howden’ ’ ( CMNC, 1 specimen; NMNH, 2 specimens) .

Pedethma weisei Lingafelter and Konstantinov , new species ( Figs. 20 , 22j , 26f)

Description. Measurements listed in Table 1. Antennae of male approximately twothirds length of body, pale at basal one­half, with apical antennomeres gradually becoming dark brown to piceous. Antennal calli slightly paler than surrounding region of head. Vertex with one setiferous pore on each side between upper portion of eyes. Supracallinal and midfrontal sulcus poorly developed, midfrontal sulcus deeper after point of bifurcation. Supraantennal sulcus well developed. Supraorbital sulcus absent. Midfrontal sulcus straight, supraantennal sulcus concave, bent around antennal socket. Lower part of antennal callus narrow and long, extending far between antennal socket and frontal ridge. A few large pores separate callus and orbit. Orbit extremely narrow, one­half times width of antennal callus. Antennal socket situated close to eye margin. Upper part of frontal ridge between antennal calli 0.63 times width of transverse diameter of antennal socket. Lower part of frontal ridge forming almost straight angle with upper part, slightly narrower than latter. No ridges situated between lower part of frontal ridge and lateral third of clypeus. Anterofrontal ridge absent. Eye 2.13 times higher than wide. Pronotum nearly rectangular with only slight antemedial bulge and posterior constriction. Posterior pronotal transverse impression slightly sinuate, short and not conspicuous (as in P. kirejtshuki ) due to the numerous, coarse punctures surrounding it. Pronotum reddish­brown with piceous lateral margins and dark brown anterior and posterior margins. Elytra ( Fig. 20 ) conspicuously maculate with piceous sutural and lateral margins and a darkened periscutellar region. Otherwise reddish­brown. Abdominal venter and metasternum piceous. Aedeagus ( Figs. 22j , 26f) relatively short with ante­apical constriction and round­ ed apex with complex pattern of medial­longitudinal impressions with interior elevations running from apex to base. Aedeagus thickened and straight from lateral perspective. Spermatheca unknown (no females known for this species).

Diagnostic and Other Comments. This species has a coarsely and heavily punctate pronotum similar to P. kirejtshuki . However, the punctures are moderately evenly spaced and not touching, in contrast to those of P. kirejtshuki which are mostly confluent. The longitudinal aedeagal excavations and impressions, the piceous margins of the pronotum and elytra, the piceous venter with pale legs, and the nearly rectangular pronotum readily distinguish this species.

Distribution. Forests of Southeast New South Wales (Map 1, locality #16).

Etymology. The specific epithet refers to the prolific early twentieth century coleopterist, J. Weise.

Material. Holotype (Male): ‘‘ AUSTRALIA: NSW, Monga State Forest , 19– 24.I.1984, L. Masner, Eucalyptus forest, 700 m.; on ferns’’ ( CMNC).

Paratypes. ‘‘[ AUSTRALIA]: Clyde Mtn., NSW, West slope, 1 Feb. 1973, D. H. Colless’ ’ ( ANIC, 2 specimens); ‘‘[ AUSTRALIA]: Clyde Mtn., NSW, West slope, 1 Feb. 1973, D. H. Colless,’’ ( NMNH, 1 specimen) .

Figures

Fig. 16. Dorsal habitus of Pedethma species. a) P. malandensis Weise; b) P. sinuatipenis Lingafelter and Konstantinov, new species; c) P. cookensis Lingafelter and Konstantinov, new species; d) P. demiensis Lingafelter and Konstantinov, new species; e) P. pinnipenis Lingafelter and Konstantinov, new species; f) P. humeromaculata Lingafelter and Konstantinov, new species; g) P. seymourensis Lingafelter and Konstantinov, new species.

Fig. 24. Aedeagi of Pedethma species. a) P. malandensis Weise, ventral view; b) P. australiensis Lingafelter and Konstantinov, new species, ventral view.

Fig. 15. Dorsal habitus of Pedethma species. a) P. howdeni Lingafelter and Konstantinov, new species; b) P. kirejtshuki Lingafelter and Konstantinov, new species; c) P. fuscipennis (dark morph); d) P. kurandensis Lingafelter and Konstantinov, new species (pale morph); e) P. kurandensis Lingafelter and Konstantinov, new species (dark morph); f) P. nigra Lingafelter and Konstantinov, new species.

Fig. 22. Aedeagi of Pedethma species. a) P. australiensis Lingafelter and Konstantinov, new species (dark morph); b) P. australiensis Lingafelter and Konstantinov, new species (pale morph); c) P. fuscipennis Weise; d) P. fuscipennis Weise; e) P. pubescens Lingafelter and Konstantinov, new species; f) P. howdeni Lingafelter and Konstantinov, new species; g) P. kirejtshuki Lingafelter and Konstantinov, new species; h) P. kurandensis Lingafelter and Konstantinov, new species; i) P. maculata Lingafelter and Konstantinov, new species; j) P. weisei Lingafelter and Konstantinov, new species; k) P. nigra Lingafelter and Konstantinov, new species.

Fig. 28. Spermathecae of Pedethma species. a) P. fuscipennis Weise (dark morph of Allyn River locality); b) P. fuscipennis Weise (pale morph of Allyn River locality); c) P. fuscipennis Weise (Dilgry River locality); d) P. howdeni Lingafelter and Konstantinov, new species; e) P. nigra Lingafelter and Konstantinov, new species; f) P. maculata Lingafelter and Konstantinov, new species; g) P. fuscipennis Weise; h) P. fuscipennis Weise; i) P. pubescens Lingafelter and Konstantinov, new species; j) P. suturalis Weise; k) P. kurandensis Lingafelter and Konstantinov, new species (dark morph); l) P. kurandensis Lingafelter and Konstantinov, new species (pale morph).

Fig. 29. Spermathecae of Pedethma species. a) P. fuscipennis Weise (dark morph of Allyn River locality); b) P. fuscipennis Weise (pale morph of Allyn River locality); c) P. nigra Lingafelter and Konstantinov, new species; d) P. maculata Lingafelter and Konstantinov, new species; e) P. pubescens Lingafelter and Konstantinov, new species; f) P. suturalis Weise.

Fig. 23. Aedeagi of Pedethma species. a) P. sinuatipenis Lingafelter and Konstantinov, new species, ventral view; b) P. sinuatipenis Lingafelter and Konstantinov, new species, lateral view; c) P. cookensis Lingafelter and Konstantinov, new species, ventral view; d) P. cookensis Lingafelter and Konstantinov, new species, oblique lateral view; e) P. demiensis Lingafelter and Konstantinov, new species, ventral view; f) P. demiensis Lingafelter and Konstantinov, new species, lateral view; g) P. pinnipenis Lingafelter and Konstantinov, new species, ventral view; h) P. humeromaculata Lingafelter and Konstantinov, new species, ventral view; i) P. humeromaculata Lingafelter and Konstantinov, new species, dorsal view.

Fig. 27. Aedeagi of Pedethma species. Scale = 0.5 mm. a) P. sinuatipenis Lingafelter and Konstantinov, new species; b) P. humeromaculata Lingafelter and Konstantinov, new species; c) P. pinnipenis Lingafelter and Konstantinov, new species; d) P. australiensis Lingafelter and Konstantinov, new species; e) P. demiensis Lingafelter and Konstantinov, new species; f) P. seymourensis Lingafelter and Konstantinov, new species.

Fig. 31. Spermathecae of Pedethma species. m) P. pinnipenis Lingafelter and Konstantinov, new species; n) P. humeromaculata Lingafelter and Konstantinov, new species; o) P. seymourensis Lingafelter and Konstantinov, new species; p) P. demiensis Lingafelter and Konstantinov, new species.

Fig. 14. Dorsal habitus of Pedethma species. a) Pedethma suturalis Weise; b) P. australiensis Lingafelter and Konstantinov, new species (dark morph); c) P. australiensis Lingafelter and Konstantinov, new species (pale morph); d) P. maculata Lingafelter and Konstantinov, new species; e) P. fuscipennis Weise (light morph); f) P. pubescens Lingafelter and Konstantinov, new species.

Fig. 21. Pedethma pubescens Lingafelter and Konstantinov, new species, dorsal habitus.

Fig. 33. Vaginal palpi in Pedethma species. a) P. kurandensis Lingafelter and Konstantinov new species; b) P. fuscipennis Weise; c) P. suturalis Weise; d) P. pubescens Lingafelter and Konstantinov new species.

Fig. 32. Spermathecal surface sculpturing in Pedethma. a) P. suturalis Lingafelter and Konstantinov, new species; b) P. howdeni Lingafelter and Konstantinov, new species.

Fig. 20. Pedethma weisei Lingafelter and Konstantinov, new species, dorsal habitus.