Paradoxichnium tumlinense, Ptaszyński & Niedźwiedzki, 2004
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publication ID |
https://doi.org/ 10.5281/zenodo.13510407 |
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persistent identifier |
https://treatment.plazi.org/id/03F387F5-163F-7C2F-FFBE-FDEC886A6061 |
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treatment provided by |
Felipe |
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scientific name |
Paradoxichnium tumlinense |
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Paradoxichnium tumlinense ichnosp. nov.
Figs. 17A, B View Fig , 18A, B View Fig , 19A–E View Fig , 20A, B View Fig , 21A, B View Fig , 29C View Fig , 35A View Fig 2 View Fig .
Holotype: Set of right pedal and manual imprints, Muz. PIG OS−220/27: 44, 45 ( Figs. 17A, B View Fig , 19C View Fig , 20A View Fig 1 View Fig ).
Type locality: Tumlin Gród quarry near the village Tumlin, 11 km NW of Kielce, Holy Cross Mts., Poland.
Type horizon: Tumlin beds, Lower Buntsandstein, uppermost Permian.
Derivation of the name: From the name of the village of Tumlin, the type locality.
Material.—Muz. PIG OS−220/27: 36–55 ten consecutive sets of imprints ( Figs. 17A, B View Fig , 18A View Fig , 19A View Fig , 20A View Fig 1 View Fig , A 2 View Fig , A 4 View Fig , 21A View Fig 1 View Fig ); the fifth of them is designed as the holotype, Muz. PIG OS−220/27: 44, 45 ( Figs. 19C View Fig , 20A View Fig 1 View Fig ); Muz. PIG OS−220/27: 139–146 four consecutive sets of pedal and manual imprints ( Figs. 19B View Fig , 21A View Fig 2 View Fig ); Muz. PIG OS−220/27: 82, 83 isolated set of imprints; Muz. PIG OS−220/39: 13–18 three consecutive sets of pedal and manual imprints ( Figs. 18B View Fig , 21B View Fig ); Muz. PIG OS−220/27: 167–178 six consecutive sets of pedal and manual imprints ( Fig. 35A View Fig 2 View Fig ).
Diagnosis.—Relatively small Paradoxichnium with a pedal imprint of about 50 mm length. Pedal pace angulation is 106–118°; that of manual ranges 151–167°. Divarication of pedal digit III from the midline is about –14°, when manual imprints are strongly inclined to the midline at about +38°. Pedal digits I–V diverge at about 40°.
Description.—Trackway. In the trackway Muz. PIG OS−220/27: 36–55 ( Figs. 17A, B View Fig , 18A View Fig , 19A View Fig , 20A View Fig 4 View Fig , 21A View Fig 1 View Fig , with the holotype, Muz. PIG OS−220/27: 44, 45) the pedal stride is 270–325 mm; that of manus attains 290–315 mm. In this trackway, interpreted as result of fast movement, manual imprints are situated on the same level as pedal ones. The pedal pace width (90–105 mm) and that of manus (15–40 mm) differ greatly, as do the respective pace angulations, 106–118° and 151–167°. This great difference is caused by an unusually narrow manual pace width. While pedal imprints diverge from the midline at –14° on average, manual imprints are inclined strongly to the midline at average angle +38°. In trackway, Muz. PIG OS−220/27: 167–178 ( Fig. 35A View Fig 2 View Fig ) interpreted as result of slow gait, manual imprints are situated slightly before pedal ones. In this trackway, pedal pace angulation reaches only 102°, and manual one 128°. The stride ranges 250–260 mm, while the pedal and manual width of pace equals 100 mm and 60 mm, respectively. The trackway of a slowly moving animal, Muz. PIG OS−220/27: 139–146 ( Figs. 19B View Fig , 21A View Fig 2 View Fig ) shows pedal and manual pace angulations 90° and 123° (respectively). Manual imprints are strongly inclined towards the midline at +40°, situated in slow gait slightly before pedal ones (Muz. PIG OS−220/27: 167–178, Muz. PIG OS−220/27: 139–146, Muz. PIG OS−220/39: 13–18, Figs. 18B View Fig , 19B View Fig , 21A View Fig 2 View Fig , B, 35A View Fig 2 View Fig ) or on the same line (Muz. PIG OS−220/27: 36–55, Figs. 17A, B View Fig , 18A View Fig , 19A View Fig , 20A View Fig 4 View Fig , 21A View Fig 1 View Fig ). Animals of similar size made all known trackways.
Pes.—Pedal imprints in trackway Muz. PIG OS−220/27: 36–55 are 50 mm long by 46 mm wide. In the digit group I–IV, 45 mm long by 38 mm wide, with greatest load on digits I–III, the digit IV is the longest. Digits I–IV diverge of 26–30°; the angle between axes of digits I and V is low, 62–64°. Digits increase in length from I to IV attaining 23, 28, 32, 34 mm (respectively). Digit V, often visible, is long as like as digit IV, 34 mm, diverging from digit I at 35–48°. Pedal imprints shows the greatest load on digits I–III. All five digits have claw marks ( Figs. 19C, E View Fig , 20A View Fig 1 –A View Fig 3 View Fig ).
Manus.—Manual imprints are 40 mm long by 30 mm wide. Digit group I–IV is 29 mm long and 26 mm wide. Digits I–IV are relatively narrow and not strongly differentiated in length, increasing from I to IV: 16, 20, 23, 27 mm, respectively. Digit I−IV have relatively small, but distinct claw marks. Digit V, 22 mm long, is much stronger imprinted than the first four digits. It is somewhat bent, enlarged on its proximal and distal end, showing distinct claw mark. It diverges from digit IV at the relatively large angle of 47°, while divarication of digits I–IV attains only 25°.
Discussion.— Paradoxichnium tumlinense ichnosp. nov. is about two times smaller than Paradoxichnium problematicum Müller, 1959 . It shows larger pace angulation on both manual and pedal imprints, larger divarication of pedal digits I–IV and I–V, much narrower manual width of pace in respect to the pedal one and a stronger inclination of the manual axes to the midline. In comparison with Paradoxichnium radeinensis Ceoloni et al., 1986 , ichnospecies questionably accounted to this ichnogenus, Paradoxichnium tumlinense ichnosp. nov. shows much narrower manual width of pace which is almost equal to pedal one in Paradoxichnium radeinensis . In contrast with the latter, Paradoxichnium tumlinense ichnosp. nov. shows pedal axes inclined outside the midline, and larger divarication of pedal and manual axes. Moreover, its pedal digit group I–IV is narrower, than that relatively short and wide in P. radeinensis .
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