Orthomorpha fluminoris Hoffman, 1977

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2011, Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 131, pp. 1-161 : 65-68

Orthomorpha fluminoris Hoffman, 1977 Figs 5556

Orthomorpha fluminoris Hoffman 1977: 701 (D).

Orthomorpha fluminoris - Golovatch 1998: 42 (D, M).

Holotype.

♂ (MHNG), Malaysia, environs of Kuala Lumpur, Batu Caves, 24.02.1975, leg. P. Strinati.

Redescription.

Length 39 mm, width of midbody pro- and metazona 2.9 and 4.4 mm, respectively. Coloration of alcohol material upon long-term preservation dark castaneous brown with contrasting yellowish paraterga; venter, legs yellownish and tip of epiproct light red-brown, antennae also light red-brown, but antenomere 6 slightly infuscate, brownish, antenomere 7 dark brown, and tip of antennae pallid (Fig. 55).

Clypeolabral region densely setose, vertex sparsely so, epicranial suture distinct. Antennae rather short, clavate (antennomere 6 broadest), extending behind body segment 2 dorsally. Head in width <collum <segment 2 <3 <4 <5-16; thereafter body gently and gradually tapering. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior; caudal corner of paraterga nearly pointed, slightly declined ventrally and continuing collum’s convexity (Fig. 55A & B); paraterga acutangular (ca 50°), slightly extending behind tergal margin, posterior edge slightly concave (Fig. 55A & B). Tegument smooth and poorly shining, prozona very finely shagreened, metazona leathery, rugulose, below paraterga microgranular. Metaterga 2-19 with two transverse rows of setae: 2+2 in anterior (pre-sulcus) row and 3+3 in posterior (postsulcus) one, all mostly abraded, but still traceable as insertion points. Tergal setae simple, moderately long, about 1/4 metatergal length. Axial line traceable. Paraterga very well developed (Fig. 55A-G), all lying below dorsum (at about 1/4 body height), slightly upturned to subhorizontal, in lateral view moderately strongly enlarged on pore-bearing segments, thinner on poreless ones; shoulders always present, broadly rounded and narrowly bordered, fused to callus; anterior edge of paraterga 2 straight, following paraterga rounded, caudal corner of all paraterga rounded, extending invariably and increasingly beyond rear tergal margin. Calluses delimited by a sulcus both dorsally and ventrally, rather broad. Posterior edge of paraterga evidently concave, especially strongly so on segments 16-19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 5-18, incomplete on segments 4 and 19, very narrow, shallow, not reaching bases of paraterga, very finely ribbed at bottom (Fig. 55A, C & amp; F). Stricture between pro- and metazona narrow and shallow, faintly beaded at bottom down to base of paraterga (Fig. 55A-F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2-4, a low and interrupted swelling supplied with a small tooth caudally on segments 5-7, retained only as small denticles on segments 8-14, thereafter missing (Fig. 55B, D & E). Epiproct (Fig. 55E-G) conical, flattened dorsoventrally, with two small apical papillae directed ventrocaudally, subtruncate at tip; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 55G) nearly semi-circular, caudal tip very broadly rounded, setiferous knobs at caudal edge very small and well-separated.

Sterna sparsely setose, without modifications; cross-impressions shallow; a pair of strongly separated, anteroventrally directed, narrowly rounded, setose cones between ♂ coxae 4 (Fig. 55H & I). A paramedian pair of small tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.5 times as long as body height, prefemora without modifications, ♂ tarsal brushes present until on legs of segment 17.

Gonopods (Fig. 56) a little more complex. Coxa long and slender, with several strong setae distodorsally. Prefemur densely setose, about half the length of femorite + “postfemoral” part. Femorite slender, slightly curved and nearly not enlarged distad, with a small, but conspicuous ventral knob near base of lamina lateralis, with a " postfemoral" part demarcated by a distinct, oblique, lateral sulcus; solenophore with a tridentate tip, middle denticle shorter than terminal tooth, but longer than a small subterminal lobule; solenomere long, flagelliform, a short tip exposed.

Remarks.

The original description is fairly complete, also illustrated in due detail ( Hoffman 1973). The above redescription, also based on the holotype, is only meant to bring it to our standard and to present additional illustrations.