Ommatius (Pygommatius) porticus, Scarbrough & Marascia, 2003

Scarbrough, Aubrey G. & Marascia, Claudio G., 2003, Revision of Ommatius Wiedemann (Diptera: Asilidae). IV. Pygommatius subgen. nov. with twenty-five Afrotropical species, Zootaxa 228 (1), pp. 1-94 : 52-54

Ommatius (Pygommatius) porticus , sp. n.

Figs. 18 , 45 , 189­196

Male. Brown. Length, body 10.7­12.0 mm; wing 7.0­ 7.8 mm. Head: Yellow to yellowishgray tomentose. Face sparse setose, setae mostly short, longer ventrally; 6 thin, brown and 4 pencil­like yellow bristles present; FHWR 1.0:10.0­1.0:10.5. Proboscis with narrow base yellow. Antenna brownish­yellow, flagellum darkest. Frons brown­yellow tomentose, brown setose. Ocellar tubercle with longest seta as long as pedicel and scape combined. Occiput with 5­6 brown postocular bristles, longest bristle about half distance toward ocellar tubercle.

Thorax: Mesonotum largely dark brown, corners yellow laterally, yellowish­brown dorsally; tomentum mostly brown, yellowish­brown to yellow laterally and posteriorly; narrow paramedial stripe boarded by thin, yellowish­brown tomentum; setae sparse, mostly brown, 4 lateral and 2 dorsocentral brown bristles present. Scutellum dark brown, yellowish­gray tomentose, sparse short pale setae and 2 brown marginal bristles present. Mediotergite black. Pleuron mostly yellow, anepisternum, katepisternite and sometimes meron black; yellow to yellowish­gray tomentum and pale setae present. Halter yellow.

Wing ( Fig. 18 ): Apical third and posterior two­thirds dense microtrichose. Cell r 4 triangular, vein R 5 slightly undulant. Apex of cell M 3 slightly curved, just beyond crossvein rm.

Leg: Coxae and anterior 2 trochanters yellow; hind trochanter brown posteriorly, 1 short brown bristle present. Femora mostly yellow; fore and middle femora yellowishbrown to brown apically; apical third­half of hind femur brown, brown sometimes extends dorsally and ventrally to basal third. Fore femur (see O. brevicornis , Fig. 23 ) ventrally with row of 6­8 yellow bristles. Middle femur with 3 long, thin, yellow bristles anteriorly; only setae present ventrally, bristles absent. Hind femur with only 4­5 anterior brown bristles; anteroventral row with brown setae, about as long as vertical height of hind femur; 1 ventrobasal bristle present; abundant setae and 3­5 bristles present posteriorly; HFWLR 1.0:3.9­1.0:4.3. Tibiae yellow with narrow apices usually brown; fore and middle tibiae with long, thin and yellow bristles, longest about three­fourths as long as fore tibia. Hind tibia ( Fig. 45 ) mostly yellow, narrow apex and base brown and brownish­yellow respectively; shaft flattened medially before preapical constriction. Tarsi with basal tarsomere of fore and middle tarsi yellow; remaining tarsomeres light brown to brown; fore tarsus laterally with 1­2 yellow bristles.

Abdomen: Mostly dark brown, wide lateral margins of tergites 1­4 and sternites 1­4 entirely yellow, with brown tomentum and brown setae dorsally; tergite 6 constricted, midapical margin of tergite 6 slightly produced, apical corner of tergite 3 with 2­3 and entire narrow lateral margin of tergite 4 with a fringe of long, yellow, sometimes brown and yellow mixed, bristles. Sternites 1­6 yellow to yellowish­brown; sternites 4­6 with abundant, erect, thin, yellow bristles; none exceptionally long or thick; sternite 7 mixed yellow and brown setose; sternite 8 brown setose.

Terminalia ( Figs. 189­193 ): Ventral lamella with long median ridge, latter with abundant, long, thin, yellow setae basally and thicker brown setae apically. Epandrium 3 branched; middle and ventral branches capitate, the former slightly longer, the latter with 1 flat, leaf­like, and 4­5 hooked bristles. Gonostylus podiform. Gonocoxite laterally with minute setae arising from large sockets. Hypandrium without apical tuft of bristles.

Female. Differs from male as follows. Length, body 9.0­ 10.5 mm; wing 7.0­ 7.8 mm. Head: Bristles thinner than in male. FHWR 1.0:10.1­1.0:10.7. Mystax with yellow or black bristles. Leg: Normal, without unusual shapes of hind femur and hind tibia and unusual vestiture of fore and hind femora.; HFWLR 1.0:5.4­1.0:5.8. Abdomen: Without unusual vestiture and constriction in males. Tergites 1­7 usually with narrow lateral margin yellow; tergites 5­8 with an apical row of bristly setae, at least 1 in apical corner of tergite 4. Sternites with mostly yellow vestiture; sternites 5­7 apically with several bristly, brown setae, and laterally with 2­3 bristles. Tergite 9 membranous dorsally, extremely short. Terminalia ( Figs. 194­196 ): Three spermathecae present; duct basal. Sternite 8 with apex acutely triangular, with medioapical ridge that apically forms a small hooked spine; surface thin, membranous; several long, thick, brown bristles present.

SPECIMENS EXAMINED. CÔTE D'IVOIRE: Holotype ♂, allotype ♀, Banco Nat. Park N. Abidjan / S. side 23­27.iv.89 / 05 o 22' N: 04 o 03' W / JGH Londt edge of wide tract / in forest ( NMSA) . Paratypes: CÔTE D'IVOIRE: 1 ♂, 9 ♀, same data as holotype ( NMSA) ; 1 ♀, Asagny Nat. Park / roadway between gate and camp / 26.iv.89 05º14’N: 04º49’W / JGH Londt forest and forest edge ( NMSA) ; 1 ♀, Haute / Cerde de Mankono / entré Kourokoro and Mankono / (F. Fleury) / A. Chevalier / vi.10 ( MNHN) ; 4 ♀, Lamto / Gal. forestière / 24.x.70 / D. Lachaise ( MNHN) ; 1 ♀, Lamto / 13.x.70 / D. Lachaise ~ forêt galerie ( MNHM) ; 1 ♀, same data except 4.x.70 ~ Lisière / forêt galerie ( MNHN) ; 1 ♀ (Fernando­Poo) Forêt / RTE de Concepcion ~ 4km. / Aprés Carref. De Moka Alt. 400 m / 13.xi . 70 L. Matile ( MNHN) ; 3 ♂, 1 ♀, Fauch Quant / 16.i.71 / GFB / D. Lachaise / ( MNHN) ; 2 ♀, Fauch Quant / GFB / 25.ix.70 ( MNHN) ; 1 ♂, same data except 3.iv.71 ~ Lamto / D. Lachaise ( MNHN) ; 1 ♀, Fauch Quant / D1 VG / 7.xii.70 ( MNHN) ; 1 ♀, Lamto Bas Fond / Savane / non brûleé / 22.i.71 / D. Lachaise ( MNHN) ; 4 ♂, 4 ♀, Marigot / Salé ~ 7.vi.71 ~ Museum Paris / Lamto / D. Lachaise ( MNHN) ; 1 ♂, same data except 19.v.71 ( MNHN) ; 1 ♀, same data except Pandanus 2 / 6.v.71 14h ( MNHN) ; 1 ♀, same data except Raphia­I / 8.v.71 ( MNHN) ; 1 ♂, same data except Zonaphia / 7.v.71 ( MNHN) ; 1 ♀, same data except J. P. Cola ( MNHN) ; 2 ♂, Marigot / Salé / 13.i.71, 15.i.71 ~ Lamto / D. Lachaise ( MNHN) ; 2 ♂, 1 ♀, Marigot / Salé ~ 26.i.71 ~, Lamto / D. Lachaise ( MNHN) ; 1 ♂, Marigot / Salé / 15.i.71 ~ Lamto / D. Lachaise ( MNHN) ; 1 ♀, Marigot / Salé 9h / 3.iii.71, Lamto / D. Lachaise ( MNHN) ; 1 ♂, same data except rn­ny / 15.ii.71 ( MNHN) ; 1 ♂, Ficus capensis / fruits à terre / Lamto 7.i.71 / D. Lachaise ( MNHN) ; 1 ♀, Savane non / brûleé / bas depente / 7.i.71 / Zone Afromomum / bosquet / pres GFB / D. Lachaise ( MNHN) ; 1 ♂, Lamto / 6.vii.71 / D. Lachaise ~ Galerie ( MNHN) ; 1 ♂, 15.viii.72 / rive du Bandama Blac / radier de la route de / Tort a à Niakara­ mandou­ / gou, 5º32’W­ 8º46’N / J. Hamon ( MNHN) ; 1 ♀, Lisière / F. Plateau / Foro­ Bouaké / 24.vii.74 / G. Couturier ~ O. R.S. T.O.M. ( MNHN) ; 1 ♀ Dabadougou / 16.xi.74 / G. Couturier, bord Marigot ~ O. R.S. T.O.M. ( MNHN) ; 1 ♀, Bouaké / Foro­Foro / 17.x.74 / G. Couturier ~ O. R.S. T.O.M. ( MNHN) ; 1 ♂, 1 ♀, same data except foret galarie / M5B ( MNHN) ; 1 ♀, Gal. For. Band. / Fauch. Quant. / 9.xi.70 ( MNHN) ; 3 ♀, Adiopodumé / 27.iii.75 / G. Couturier ( MNHN) ; 1 ♂, 2 ♀, same data except 1.xi.74, 7.iii.75, 4.iv.75 / G. Couturier Chasse, ( MNHN) ; 1 ♀, same data except Chasse / 23.i.75 ( MNHN) ; 1 ♀, Cascade / Mam / 21.xi.74/ G. Couturier ( MNHN) ; 2 ♀, Taï / 15­20.viii.78 / G. Couturier ~ forêt dense / sempervirente ( MNHN) . GUINEA: 1 ♂, Camp 4 (1000 m), Museum Paris / Nimba (Guinee) / M. Lamette 11.vi.48 ( MNHN) . NIGERIA: 1 ♀, Lagos, G. Strachan ( BMNH) ; 1 ♂, 1 ♀, T. Cacao ~ C. R.I.N. / 7:ll:1986 / Gambari / R. H. Booker ( BMNH) ; 1 ♀, Ikeja Airport / Lagos Nigeria / 23.xii.48 / B. Malkin ( CASC) .

Distribution. A forest edge species captured throughout the year in West Africa.

Etymology. Latin, porticus , for ‘gallery or path’ [forêt galerie] in a forest.

Remarks. In addition to the characters in the key, the male O. porticus is recognized by the combined characters of the terminalia ( Figs. 189­193 ) and the two rows of erect setae on sternites 4­6.

Figures

FIGURES 11­20. Right wing of male Ommatius (Pygommatius) spp. 11. O. imaginus, sp. n..; 12. O. limbus, sp. n..; 13. O. litoreus sp. n.; 14. O. lulua, sp. n.; 15. O. magnipes, sp. n.; 16. O. ornatipes Engel; 17. O. pectinus sp. n.; 18. O. porticus, sp. n.; 19. O. renudus, sp. n.; 20. O. strigiatus, sp. n.

FIGURES 39­48. Left hind tibia of male Ommatius (Pygommatius) spp. 39. O. comosus, sp. n.; 40. O. fluvius, sp. n.; 41. O. imaginus, sp. n.; 42. O. limbus, sp. n., anterior; 43. O. litoreus, sp. n.; 44. O. neglectus Bromley; 45. O. porticus, sp. n.; 46. O. renudus, sp. n.; 47. O. strigiatus, sp. n.; 48. O. vultus, sp. n., inner surface.

FIGURES 189­196. Male terminalia of Ommatius porticus, sp. n. 189­191. Lateral, dorsal and ventral views; 192. Gonostylus; 193. Aedeagus, lateral view. Female terminalia. 194. Spermatheca; 195. Genital fork; 196. Sternite 8, ventral view.

FIGURES 21­28. Right wing of male Ommatius (Pygommatius) spp. 21. O. talus, sp. n.; 22. O. vultus, sp. n.; 23­28. Right fore femur of male Ommatius (Pygommatius) spp. showing arrangement of anteroventral bristles. 23. O. brevicornis Curran; 24. O. grossus, sp. n.; 25. O. limbus, sp. n.; 26. O. litoreus, sp. n.; 27. O. strigiatus, sp. n..; 28. O. vultus, sp. n.