Ogyges ratcliffei Cano, 2014

Cano, Enio B., 2014, Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly, Zootaxa 3889 (4), pp. 451-484 : 471-473

Ogyges ratcliffei Cano new species

Figs. 15–16 , 21v

Diagnosis. Ogyges ratcliffei seems most similar to O. nahuali and O. cavei . The smaller size, the noticeable blue reflections on brilliant areas, the narrow and flat lamellae and the shagreened metasternal marginal groove, separate O. ratcliffei from O. cavei . The lack of punctures surrounding metasternal disc and central horn directed upward, distinguish the new species from O. nahuali .

Description. Holotype, female, black adult with blue iridescence. Total length 33.02, elytral length 17.9, pronotal length 8.21, pronotal width 11.95, humeral width 11.12. Head: anterior border of labrum slightly concave. Clypeus slightly inclined, transversaly convex, anterior border straight and thick, with very small central invagination and with external anterior angles acute; separated from the mediofrontal area by a rugose sink running between the conical and moderate mediofrontal tubercles. Mediofrontal area smooth, without internal tubercles and posterofrontal ridges. A small, bare, smooth fossa present in front of the mediopostfrontal structure. Lateropostfrontal areas glabrous and smooth. Central horn moderately long, with apex directed upward and forward, with unclear median longitudinal groove on posterior 1/3; lateroposterior tubercles short, rounded, curved backward and separated from central horn. Postfrontal groove smooth and shallow, deeper at sides. Supraorbital ridge with equal anterior tubercles; posterior 1/2 not bifurcate with supraorbital fossae small; external ridge marked. Ocular canthus with apex straight covering half of one eye. Eyes reduced. Eye width = 0.61 mm. Interocular width = 6.61 mm. Head width (measured between tips of canthi) = 8.06 mm. Ratio width of both eyes/ head = 0.15. Postorbital pits located between supraorbital ridge and eye, punctate setose. Ligula slightly protuberant basally, with apical central tooth eroded, anterior ventral carina absent, setose punctures on the median area. Lateral lobes of mentum with abundant setose punctures; medial basal mentum glabrous (except two punctures on one side near the lateral scar); lateral basal scars elongate to external side, punctate-setose and brilliant. Hypostomal process elongate, with poorly marked lateral depression, wide medially and narrow in the apical third. Infraocular ridge present, short and smooth, proximal area declivous and setose, distal area punctatesetose. Mandible with dorsal tooth occupying 1/2 of his length; internal face in dorsal view slightly opaque, not granular. Antennal club not concave, penultimate antennomere wider than the antepenultimate ( Figs. 15–16 ).

Thorax: Lateral fossa of pronotum without punctures except minute striate-punctures ahead and behind, visible at high magnification. Pronotum with marginal groove narrow and smooth, with minute striate-punctures visible at high magnification; anterior angles rounded. Prosternellum opaque on anterior half (or slightly more) and shiny on posterior half. Mesosternum with lateral depressions elongate and rugose (shagreened). Mesepisternum with elongate rugose area (shagreened), more marked and oval apically. Metasternum anterior angles without setae, apical borders shagreened; disc without punctures; marginal groove glabrous, narrow and shagreened throughout, posteriorly two times wider than medially.

Elytra: Shiny, with blue reflections that give a dull appearance. Striations well marked; with minute, superficial punctures on striae 1–4, gradually deeper and wider between 5–10 and well marked from 1–10 on posterior 1/3; junctions of striae 1 and 10 with sparse extra punctures. Anterior border of elytra vertical, with a few minute setae on intrestriae 3–6.

Legs: Profemur with anterior-ventral groove marked; metafemur moderately widened; mesotibia with one spine.

Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.

Material examined. 1 specimen.

Type material. Holotype deposited at UVGC: HONDURAS: Olancho, La Picucha, 11 km N Catacamas, 14.92740, -85.90983. Bosque nuboso, 1800–2100 m, 8–14 V 2010. L. Sáenz (LSD 451).

Etymology. Named after the Team Scarab leader, Brett C. Ratcliffe, honoring his studies on the dynastine beetles of the world.

Distribution. The species is known only from the holotype collected in cloud forest between 1800–2100 m, in Olancho, Reserva La Picucha, Honduras ( Fig. 4 ).

Figures

FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi.

FIGURE 15. Holotype of Ogyges ratcliffei: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view.

FIGURE 16. Dorsal aspect of head of Ogyges ratcliffei.

FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili.