Ogyges menchuae Cano, 2014

Cano, Enio B., 2014, Ogyges Kaup, a flightless genus of Passalidae (Coleoptera) from Mesoamerica: nine new species, a key to identify species, and a novel character to support its monophyly, Zootaxa 3889 (4), pp. 451-484 : 465-467

Ogyges menchuae Cano , new species

Figs. 11–12 , 21p

Diagnosis. Ogyges menchuae is most similar to O. cakchiqueli Schuster & Reyes-Castillo , O. championi (Bates) , and O. kekchii Schuster & Reyes-Castillo. These species are in the “ O. laevior ” [= O. championi ] lineage of Schuster & Reyes Castillo (1990). The large internal tubercles separate it from similar species, except from O. tzutuhili Schuster & Reyes-Castillo and O. marilucasae Reyes-Castillo & Castillo , but is easily separated by the narrow antennal club (very wide in O. tzutuhili and O. marilucasae ), and the smaller size.

Description. Holotype male, black adult. Total length 29.24, elytral length 16.20, pronotal length 8.38, pronotal width 10.61, humeral width 10.03. Head: anterior border of labrum straight, dorsal surface granular. Clypeus vertical, anterior border straight, posteriorly microgranulate, separated from the mediofrontal area by an inflection; mediofrontal tubercles conical, with apex rounded, moderately enlarged and markedly visible in lateral view. Mediofrontal area slightly microtuberculate, moderately roughened; internal tubercles present and well developed, directed forward and slightly upward, connected to the curved and strongly marked posterofrontal ridges; angle of posterofrontal ridges concave and united to the central horn. Laterofrontal areas roughened, with sparse granulations. Lateropostfrontal areas glabrous, smooth, somewhat roughened and with sparse microtubercles or granulations. Central horn short with apex conical, not free and with median longitudinal (mostly deep) groove dorsally in basal half; lateroposterior tubercles elongate, rounded, directed forward, separated from the central horn by a small groove with sparse or moderate microgranulations. Postfrontal groove deeper at sides and separated from lateropostfrontal areas. Supraorbital ridge with mostly unequal or almost equal anterior tubercles; posterior 1/2 bifurcate, with the supraorbital fossae very deep; external ridge well marked. Ocular canthus with apex swollen, apex forming a right angle. Eyes reduced, width = 0.54 mm (each eye). Interocular width = 6.15 mm. Head width (measured between tips of canthi) = 6.92 mm; ratio both eyes/head = 0.16. Postorbital pits situated immediately behind supraorbital ridges, almost rectangular, shallow and with a few punctures; external borders with setose punctures. Ligula slightly protuberant basally; with a central apical tooth small and with a transversal, well marked anterior ventral ridge; setose punctures present medially. Lateral lobes of mentum with abundant setose punctures; lateral basal scars oval, brilliant and punctate setose, internal border longitudinally striate. Medial basal mentum bare and with sparse micropunctures visible at great magnification. Hypostomal process elongate, wide medially and narrow in apical third, without lateral depression. Infraocular ridge short, external and basal area punctate-pubescent. Dorsal tooth of mandible occupies 1/2 length of the mandible; internal face of mandible in dorsal view, granular. Antennal club ( Figs. 11–12 ) 1.88 mm long, 1.31 mm wide; with last two antennomeres wider than the antepenultimate, this slightly more than twice as wide as long.

Thorax: Lateral fossae of pronotum with micropunctures of rugose aspect, visible with moderate magnification. Pronotum with marginal groove narrow, widened at anterior margin; anterior angles rounded, slightly projected anteriorly immediately behind eyes. Pronotum with dense and shallow micropunctures of rugose aspect, visible at moderate magnification, running from the lateral fossae to the anterior border, internal to marginal groove; disc with sparse micropunctures visible at great magnification. Hypomeron with abundant small shallow punctures. Prosternellum completely shagreened. Mesosternum bare, lateral depressions elongate, rugose (shagreened). Mesepisternum with large and oval rugose (shagreened) area extended to posterior margin. Metasternum with anterior angles bare; disc not delimited by punctures; marginal groove bare and narrow, posterior part 1.5–2.0 times wider than the median part.

Elytra: Striations marked and with small but defined punctures, wider and deeper from 5–10; interstriae 1–7 with sparse micropunctures, more abundant and visible at moderate magnification on interstriae 8–10; junctions of striations 1 and 10 without extra punctures.

Legs: Profemur with anterioventral groove marked, metafemur widened; mesotibia with two (rarely three) spines.

Abdomen: Marginal groove of sternite VII incomplete, occupies 3/5 of sternite.

Aedeagus: In ventral view phallus globose; parameres and phallobase separated; in dorsal view, ventrodorsal basal sclerotizations of the phallus present ( Fig. 12 ).

Variation (n = 20): Total length 29.36–33.15 ( = 31.43), elytral length 16.11–18.66 ( = 17.29), pronotal length 7.40–9.13 ( = 8.56), pronotal width 10.12–11.72 ( = 10.93), humeral width 9.46–10.71 ( = 10.17). Mediofrontal area more or less smooth to somewhat or moderately roughened, with scarce to barely visible microgranulations. Some specimens present an anterior keel in the concavity of the angle of posterofrontal ridges.

Material examined. 35 specimens.

Type material. Holotype: GUATEMALA: Quiché, Uspantán, aldea Laj Chimel , montaña al norte de la aldea, 2100 m., VII 1998, bosque nuboso. E.B. Cano collector.

Paratypes: GUATEMALA: same data as holotype (1 male, 7 unknown) ; same data except Norte Laj Chimel , San Pedro, 2100m (1 male, 1 unknown) ; same data except Aldea Laj Chimel, A. González-Madrid, 30 V 2011 (13) ; same data except, Aldea Laj Chimel, road to San Pablo, 30 IV 2011, 15 o 27'30.69''N, 90 o 46'26.11''W, 2035 m, A. Gónzález-Madrid 30 V 2011 (3 male, 1 unknown) ; same data except, Aldea Laj Chimel, road to San Pablo, 30 IV 2011, 15 o 27'30.69''N, 90 o 46'26.11''W, 2035 m (6) ; same data except, Montaña El Amay, in logs 15 VI 2012, A. Zamora (2 males) .

Holotype deposited in UVGC . Paratypes deposited in UVGC, IEXA, USAC, IBUNAM, SNM , and MNHN.

Etymology. Named in honor of the Nobel Prize winner, Dr. Rigoberta Menchú, who was born in Laj Chimel. Distribution and ecology. Known only from middle altitude cloud forest at Cerro El Amay, of Laj Chimel,

Quiché, Guatemala, 1800–2100 m altitude ( Fig. 4 ) .

Figures

FIGURE 4. Map of northern Central America showing the locality of the new species. Light blue square, Ogyges menchuae; rose circle, O. handali; blue rhombus, O. cavei; yellow triangle, O. llama, O. mutenroshii; purple cross, O. toriyamai; red star, O. laurae, O. ratcliffei, and light green circle, O. sandinoi.

FIGURE 11. Paratype of Ogyges menchuae: a. lateral view of head. b. dorsal view. c. ventral view. d. lateral view.

FIGURE 12. Ogyges menchuae: a. dorsal aspect of head. b. ventral view of aedeagus. c. lateral view of aedeagus. d. dorsal view of aedeagus.

FIGURE 21. Details of suprainternal teeth of right mandibles (left in O. cavei) of the 25 known species of Ogyges: a. O. adamsi. b. O. aluxi. c. O. cakchiqueli. d. O. cavei. e. O. championi. f. O. coxchicopi. g. O. crassulus. h. O. furcillatus. i. O. handali. j. O. hondurensis. k. O. kekchii. l. O. laevissimus. m. O. laurae. n. O. llama. o. O. marilucasae. p. O. menchuae. q. O. monzoni. r. O. mutenroshii. s. O. nahuali. t. O. politus. u. O. quichensis. v. O. ratcliffei. w. O. sandinoi. x. O. toriyamai. y. O. tzutuhili.