Nectamia savayensis ( Günther, 1872 )

Fraser, Thomas H., 2008, Cardinalfishes of the genus Nectamia (Apogonidae, Perciformes) from the Indo-Pacific region with descriptions of four new species, Zootaxa 1691 (1), pp. 1-52 : 32-37

Nectamia savayensis ( Günther, 1872)

Figures 6F , 13 , 18 and Tables 2–3, 8

Type material: Holotype Apogon savayensis BMNH 1871.9.13.218; 48 mm SL; Savai’i (Savay, Western Samoa); Godeffroy Coll.

Additional Material: Indian Ocean: Mozambique: SAIAB 3311 ; (2, 47, 53); Island das Rolas ; 11 Aug 1951 ; x-ray. SAIAB 3318 ; (1, 58); Pinda Reef; 9 Sept. 1956 ; x-ray. SAIAB 3321 ; (1, 42); Mozambique, Pinda Reef , June 1951 ; x-ray. SAIAB 3315 ; (2, 36–55); Pinda Reef ; July 1950 ; x-ray. SAIAB 3317 ; (1, 65); Matemo I.; 13 Aug 1951 ; x-ray. Zanzibar: SAIAB 3320 ; (1, 56); 3 Sept 1952 ; x-ray. Madagascar; UMMZ 185908 ; (1, 44); Tulear; 11–13 July 1964 ; JB Sta. 64-22. Aldabra Islands : USNM 213130 ; (5, 31–49) Lagoon inside SE portion of West I (Ile Picard); 3 Dec. 1964 ; IIOE Cr. 9, HAF-16 . SAIAB 3314 ; (1, 68); Nov 1954 ; x-ray.

Farquhar Islands: USNM 213131 ; (21, 50–67) 6 Dec 1964; IIOE Cr 9, R . Suttkus-37; USNM 213132 ; (74, 24–62) Lagoon W. of North I; 6 Dec 1964; IIOE Cr. 9; HAF-18. Comoro Islands: USNM 213133 ; (6, 56–68) Grande Comore I; W. side edge of fringing reef; 27 Nov 1964; IIOE Cr 9, HAF-14. USNM 213134 ; (23, 27– 60) ; Lagoon ca 3.5 mi. SW of S Pt of Pamanzi I.; 24 Nov 1964; IIOE Cr. 9; HAF-9. Amirante Isles : ANSP 152769 ; (4, 39–71) D’Arros I; 5 Mar 1964; Sta. F-88. USNM 213135 ; (50, 25–67) St. Joseph Is , Ressource I; 8 Dec 1964; IIOE Cr 9, H.A. Fehlmann-19 ANSP 152792 ; (1, 14) D’Arros I; 5 Mar 1964; Sta. F-87. ANSP 164655 ; (1, 14) ; off N. end of North I; 3 Mar. 1964; IIOE-ANSP; Sta. F-78. USNM 213136 ; (1, 17) ; D’Arros I; 8 Dec 1964; IIOE Cr 9, Suttkus-40. Seychelles: Mahé I.: ANSP 152775 ; (2, 32–39) ; 19 Mar 1964; Sta. F- 119. ANSP 152791 ; (7, 41–67) Beau Vallon Bay; 15 Mar 1964. ANSP 152773 ; (6, 36–70) ; North West Bay; 9 Feb 1964; Sta. F-36. ANSP 152796 ; (16, 29–74) ; 31 Jan –2 Feb 1964; Sta. F-13. ANSP 152800 ; (5, 22–36) Souris I.; 16 Feb 1964; Sta. F-53. CAS 58762; (2, 68–73) ; Te Vega; 27 Oct 1964. SAIAB 3322 ; (1, 57) ; 16 Sept 1954. SAIAB 3319 ; (2, 64–65) ; 27 Sept 1954. BPBM 35518 ; (1, 19) ; 24 Dec 1992. Beacon I.: ANSP 152801 ; (3, 29–45) ; 25 Jan 1964; IIOE-ANSP; Sta. F-5. ANSP 152802 ; (2, 28–39) ; 3 Feb 1964; Sta. F-18. ANSP 152772 ; (1, 18) ; Off Port Victoria I.; 6 Feb 1964; Sta. F-26. ANSP 152799 ; (2, 35–69) ; 27 Jan 1964; Sta. F-7. ANSP 152798 ; (5, 30–73) ; N. of Anonyme I, Mahé vicinity; 10 Feb 1964; Sta. F-37. ANSP 152797 ; (5, 22–62) W. of NW Anonyme I., near Mahé I; 11 Feb 1964; Sta. F-44. Sta. F-114. ANSP 152771 ; (2, 24–39) Just N. Anonyme I., near Mahé I; 2 Feb 1969; Sta. F-17. ANSP 152805 ; (14, 27–63) Between Praslin & Round Is.; 22 Feb 1964; Sta. F-59. ANSP 152804 ; (1, 38) S of Round I, near Praslin I; 22 Feb 1964; Sta. F-61. ANSP 152458 ; (1, 14) ; Curieuse Island ; F-64; 23 Feb 1964. ANSP 152770 ; (11, 34–40) SE Rouge Pt, Curieuse I; 23 Feb 1964; Sta. F-62. ANSP 152803 ; (6, 35–60) ; S. of Rouge Pt, Curieuse I.; 24 Feb 1964; Sta. F-66. ANSP 152774 ; (9, 20–65) Faon I; 24 Jan. 1954; Sta. F-3. CAS 35363; (23, 53–67) ; Cerf I.; Te Vega Cr. 5; 21 Oct 1964. CAS 35369; (15, 55–76) ; St. Anne I.; Te Vega Cr. 5; 19 Oct 1964. SAIAB 3312 ; (1, 58) Silhouette I; 2 Oct 1954; x-ray. Reunion: USNM 345778 ; (1, 34) ; west coast near St Leu; 7 Feb 1997. Mauritius: FMNH 75711 ; (57, 42–74) ; Black Rock Point; 24 May 1964; LPW-41. USNM 349771 ; (11, 35–78) ; M28; 16 May 1995. USNM 213137 ; (20. 43–77) ; reef off Port Louis; 12 June 1964; IIOE Cr. 6. USNM 213138 ; (1, 73) ; reef off Port Louis; 11 June 1964; IIOE Cr. 6. SAIAB 2120 ; (4, 40–92) ; La Morne Brabant; THF-SA-28; 7 Mar 1971; x-ray. SAIAB 2311 ; (11, 33–38) ; Palamar; THF-SA-44; 27 Mar. 1971; x-ray. SAIAB 2234 ; (9, 44–52) ; near Riambel; THF-SA-42; 26 Mar. 1971; x-ray. BPBM 20102 ; (3, 46–67) ;Trou d’Eau Douce; 31 Oct. 1973. St. Brandon Shoals : SAIAB 1893 ; (3, 66–71) ; Tortue I; 15 Mar 1971; THF SA- 35. SAIAB 2207 ; (1, 44) ; Raphael I.; 18 Mar 1971; THF SA-36. Chagos Archipelago: Diego Garcia Atoll: USNM 213139 ; (2, 26– 55) ; 7 July 1967; HAF 67-36. USNM 213140 ; (2, 31–62) ; 26 June 1967; HAF 67-18. USNM 213141 ; (18, 26–59) ; 13 June 1967; HAF 67-5. USNM 213142 ; (10, 30–54) ; 12 June 1967; HAF 67- 3. USNM 213143 ; (2, 32–67) ; 9 July 1967; HAF 67-38. USNM 213144 ; (3, 29–37) ; 20 July 1967; HAF 67- 49. USNM 213145 ; (2, 47–52) ; 18 June 1967; HAF 67-11. Maldive Islands: FMNH 75707 ; (14, 22–38) Addu Atoll; 10 May 1964; LPW-40. FMNH 75670 ; (4, 32–52) Tiladummati Atoll; 31 March 1964; LPW-29. FMNH 75683 ; (2, 41–44) Ari Atoll; 22 Apr 1964; LPW-35. FMNH 75652 ; (2, 66–67) S. Malé Atoll; 18 Mar 1964; LPW-24. FMNH 75695 (19, 22–59) S. Nilandu Atoll; 24 Apr 1964; LPW-37. FMNH 75665 ; (77, 25– 62) Miladumadulu; 27 Mar. 1964; LPW-28. FMNH 75698 ; (115, 21–65) ; Addu Atoll 30 Apr 1964; LPW-38. BPBM 33049 ; (1, 62) ; North Malé Atoll, W. side of Huraa Island ; depth 20– 22m.; 24 Mar. 1988. SMF 4750 ; (7, 28–40) ; Ari Atoll , 3°15´N, 73°00´E; 17 Feb 1958. SMF 4761 ; (2, 43–46) ; Fadiffulu Atoll , 3°15´N, 73°00´E; 13 Apr 1958. Sri Lanka: USNM 213146 ; (1, 59) ; Trincomalee; 4 Apr 1970; CCK 69-136. USNM 213147 ; (1, 42) ; Hikkaduwa, 1¼ mi. off H Kralla; 17 Feb 1970; CCK 69-109. USNM 213148 ; (1, 64) ; Korali Pattu, Batticaloa District ; 6 June 1970; TI 70-344 . USNM 213149 ; (1, 15) ; Trincomalee; 8 Apr 1970; CCK 69-143. USNM 213150 ; (1, 22) ; Trincomalee; 27 June 1969; WSV 69-120. India: FMNH 75638 ; (3, 20–29) ; Mandapan Camp; 31 Jan 1964; LPW-15. FMNH 75627 ; (3, 68–82) ; Gulf of Mannar , Musal tivu; 27 Jan 1964; LPW-13. FMNH 75616 ; (9, 28–73) ; Gulf of Mannar , Musal tivu; 7 Jan 1964; L.P. Woods - 5. FMNH 75605 ; (9, 62–66) ; Gulf of Mannar , Musal tivu; 30 Dec 1963; LPW-3. Nicobar Is.: SMF 7759 ; (2, 38–54) ; Tillanchong I., 8°00'N, 93°30'E; 3 Sep 1958 . Cocos-Keeling Islands: ANSP 128377 ; (1, 22) ; North Keeling I; 6 Mar 1974; Sta. 22. ANSP 128374 ; (4, 20–22) West I; 28 Feb 1974; Sta. 12. ANSP 128372 ; (1, 30) ; West I; 25 Feb 1974; Sta. 6. ANSP 128376 ; (1, 27) ; Prison I; 7 Mar 1974; Sta. 23. ANSP 128373 ; (1, 16–17) ; Direction Island ; Sta 9; 27 Feb 1974. ANSP 128375 ; (1, 15–16) ; Horsburgh Island ; Sta 5; 25 Feb 1974. Thailand: ROM 70484 ; 12(16–89); Phuket, 7°55'15"N 98°16'16"E; RW 93-8; 13 Nov 1993; 8– 13 m. ROM 71478 ; 40(44–74); Phuket, 7°47'10"N 98°23'45"E; RW 93-3; 10 Nov 1993; 0– 9 m. ROM 71479 ; 20(33–66); Phuket, 7°45'40"N 98°20'2"E; RW93-4; 10 Nov 1993; 0– 6 m. ROM 72174 ; 4(12–58); Phuket, 7°73'15"N 98°22'50"; RW 93-12; 16 Nov 1993; 3– 8 m. ROM 72176 ; 1(54); Phuket, 7°72'15"N 98°19'35"E; RW 93-17; 18 Nov 1993; 15– 22 m. ROM 72177; 9(38–62); Phuket, 7°72'15"N 98°19'35"E; RW 93-18; 18 Nov 1993; 1– 9 m. ROM 72179 ; 1(62.5); Phuket, 8°83'99"N 97°39'99"E; RW 93-34; 14 Dec 1993; 0– 10 m. USNM 213152 ; (1, 33) ; Paton Bay , Phuket; 22 Mar 1963; IIOE Cr. I. Indonesia: CAS 95833; (1, 57) ; Pulau Pandan, off Padang, Sumatra; 25–26 m; 17 Apr 1997. Australia: Timor Sea: CAS 52558; (29, 24–45) ; Ashmore I.; 3 Jan. 1973. Pacific Ocean : Thailand: CAS 213345 ; GVF 2172 ; (1, 53) ; Khorn Aho Bay , 12°37'37"N 101°20'31"E; 24 Apr 1960. CAS 84058; GVF 2651 ; (1, 72) ; 1.5 mi. ESE of Town of Prachuap Khiri Khan; 19 June 1961. Viet Nam: CAS 83817; GVF 2113 ; (10, 59–70) ; Ilot du Sud , 10°29'15"N; 108°57'30"E; 10 Mar 1960; MV Stranger, Sta. 60- 96. ROM 79172; (6, 19–79) ; RW02-21; 9 Jun 2003 Singapore: BPBM 21854 ; (1, 50) ; Sulu I.; 24 July 1977. Indonesia: CAS 62476; (1, 43) ; Sangi Siang I.; JEM 87-7; 24 Sep 1987. Java Sea : BPBM 18583 ; (1, 49) ; Capelan Seribu, Pulau Petri; black and white; 17 Feb. 1975. Banda Sea : USNM 213113 ; (8, 43–73) ; SE Naira I, Banda I.; 8 Mar 1974. VGS 74-9; USNM 213187 ; (10, 32–69) ; Ambon; 14 Mar 1974; VGS 74-13. USNM 171388 (A.23359); (1, 47) ; Bauru I; 9 Dec. 1909; Molucca Sea: Sulawesi: USNM 112234 (A.23394); (1, 63) Gulf of Tomiki ; 16 Nov 1909. USNM 112190 (A.23974); (1, 49) Limbs Str.; 11 Nov 1909. Sulawesi Sea : BPBM 36709 ; (1, 67) ; Sulawesi, Maenad, Bunaken; 8m; 28 Oct 1991. Philippines: USNM 344522 ; (15, 32– 69) ; Panay, Lawigan, Tiolas ; JTW 95-4. USNM 261148 ; (9, 48–60) ; Negros I., south tip; SP 78-9; 12 May 1978. USMN 261151 ; (26, 21–62) ; Palawan Is., Bararin I., SP 78-20; 23 May 1978. USNM 261147 ; (2, 65– 71) ; Pamilikan I.; SP 78-43; 12 June 1978. USNM 329215 ; (38, 28–62) ; Negros Oriental, Siquijor I.; LK 79- 15, 15 May 1979. Sulawesi Sea : USNM 171399 (A.24005); (1, 56) ; Sitanki Reef, S. Tawi Tawi Group; 24 Sept 1909; Sulu Sea : ANSP 98137 ; (15, 34–52) ; Sibutu Island ; 27 Apr 1926. USNM 171403 (A.21558); (1, 52) ; Pilas I, S of Zamboanga ; 12 Sept 1909; USNM 171405 (A.15565); (1, 40) ; Bugsuk I, N. Balabac Str.; 5 Jan 1909. Philippine Sea : USNM 112164 (A.23655); (1, 54) ; Batan I; 8 June 1909; x-ray. China: CAS 83800; GVF 1750 ; (8, 51–64) ; Pratas Reef; 25 May 1958; Sta. HK 73. Taiwan: USNM 213155 ; (4, 49–60) ; Bay between K’en-ting and Ta-yuan Shan; 1 May 1968; VGS 68-12. USNM 213156 ; (3, 65–72) ; Off Ch’uanfan-shih, SW shore; 3 May 1968; VGS 68-14. USNM 213157 ; (13, 44–82) ; Ch’uan-fan-shih; 23 Apr 1968; VGS 68-2. BPBM 23300 ; (4, 37–83) ; S end at Ch’uan-fan-shih; 0–6m; 16 Jul 1978. South Korea: CAS 29963; (1, 18) ; Inchon area; 6–7 Jun 1973. Carolina Islands : Palau: BPBM 7225 ; (5, 37–54) ; SE of Auluptagel I.; 26 Sep. 1966. BPBM 8091 ; (6, 35–55) ; SW of Koror I.; 12 Jun 1968. CAS 83820; GVF 908 ; (1, 52) ; Ngadarak Reef; 25 Sept. 1956, Sta. 122. CAS 84055; GVF 925 ; (7, 29–61) ; Ngaruangl Reef; 5 Oct 1958; Sta. 139. CAS 83778; GVF 804 ; (3, 50–68) ; Ngadarak Reef; 12 Jul 1956. CAS 84061; GVF 897 ; (1, 74) ; Ngadarak Reef; 12 Sep 1956. CAS 84085; GVF 1385 , (3, 54–64) ; Ngadarak Reef; 11 Sep 1957. CAS 83790; GVF 490 , (2, 64–68) ; Urukthapel I.; 23 Sep 1954. CAS 84084; GVF 1397 ; (27, 17–61) ; Babelthuap I.; 22 Sep 1957. CAS 83791; GVF 1958 ; (3, 53–67) ; Ngethil I.; 13 Jul 1959. Ifaluk Atoll: CAS 83794; GVF 129 ; (22, 32–62) ; 21 Sept 1953; Sta. 8. CAS 83809; GVF 216 ; (17, 37–54) ; 20 Oct 1953; Sta. 95. CAS 83810; GVF 262 ; (21, 41–54) ; 28 Oct 1953; Sta. 141. CAS 83777; GVF 976 ; (10, 38–61) ; Ulitki Atoll; Sorlen I, W. side; 22 Sep 1956. CAS 83880; GVF 215 ; (13, 39–44) ; 20 Oct 1953; Sta. 94. Truk: BPBM 9313 ; (13, 14–56) ; 10 Jul 1969 l. Pohnpei: CAS 83788; GVF 496 ; (13, 23–53) ; 6 Oct. 1954; Sta. 193. CAS 83784; GVF 493 ; (1, 72) ; 4 Oct 1954; Sta. 190; Kapingamarangi Atoll: CAS 83798; GVF 317 ; (3, 31–42) ; Tokoteihi; 1 July 1954; Sta. 14. CAS 83970; GVF 329 ; (1, 45) ; 8 July 1954, Sta. 26. Sorol Atoll: CAS 83781; GVF 993 ; (1, 45) ; 28 Oct 1956. Mariana Islands: Guam: ANSP 85880 ; (1, 82) ; AS 00-G-6-1; 1 Jun 1900. CAS 84065; GVF 788 ; (25, 51–65) ; 14 Jun 1956. CAS 83877; (3, 40–47) ; GVF 1839 ; 10 Oct 1958; sta 11. CAS 213343 ; GVF 788 ; (1, 62.9) ; 14 Jun 1956. UGM 1418 ; (3, 53–69) ; 5 Oct 1968; GFW-111. UGM 1415 ; 7(45–71); 28 June 1968; GFW-119; 27– 33m. Saipan ANSP 164649 ; (1, 66) ; SAI-1-25; 18 Oct 1953. CAS 83894; GVF 790 ; (4, 36– 63) ; 17 June 1956; Sta. 4. CAS 83896; (9, 16–35) ; GVF 789 ; 16 Jun 1956. CAS 213326 ; (1, 56) ; G90-8; 24 Feb 1990. CAS 80770; (3, 47–51) ; G90-6; 25 Feb 1990. Yap I.: CAS 84155; GVF 1935 ; (1, 53) ; 6 Jan. 1959; Sta. 112. Marshall Islands : Arno Atoll: USNM 166599 ; (1, 29) ; Eonëb-je I.; 3 July 1950. Bikini Atoll: USNM 142415 ; (9, 19–32) ; Ari I.; 7 Aug 1946; LPS, Sta. S-46-308. Enewetak Atoll: BPBM 28971 ; (1, 51) ; Enewetak Atoll; 23m; 24–25 Sep. 1982; x-ray. BPBM 31233 ; (1, 51) ; 19 Apr 1976. Kwajalein Atoll: USNM 142428 ; (1, 19) ; near Ennylabegan I.; 1 Sept 1946; LPS, Sta. S-46-397. Rongerik Atoll: USNM 142417 ; (2, 27–28) ; Latoback I.; 14 Aug 1947; LPS, Sta. S-1041.6. Rongelap Atoll: USNM 142414 ; (27, 23–61) ; Kieshiechi I; 24 July 1946; LPS, S-46-285. Wake I.: BPBM12041 ; (1, 49) ; May 1957. SU 50057 ; (2, 47–48) . SU 51220 ; (24, 39–51) ; Between Wilkes and Peale Is; 20 Sept 1957. Papua New Guinea : Bougainville I.: USNM 268266 ; (79, 31–67) ; TeVega Cr. 6; 19 Oct. 1984. USNM 262489 ; (9, 17–54) ; Hermit Is., Amot I.; VGS 78-10; 30 Oct. 1978. USNM 213251 ; (2, 19–31) ; Kranket I.; BBC-1498; 31 May 1970. USNM 296755 (1, 47) and CAS 65301 (1, 44) ; Wongat I.; SGP 87-29; 20 May 1987. CAS 65299; (1, 54) ; Wongat I.; SGP 87- 33; 22 May 1987. Nagada Harbour : USNM 296845 ; (1, 38) and CAS 65308 (2, 44–52) ; SGP 87-23; 16 May 1987. USNM 296742 ; (1, 48) and CAS 65300; (1, 34–50) ; SGP 87-30; 20 May 1987. CAS 65302; (1, 44) ; SGP 87-19; 12 May 1987. CAS 65305; (1, 46) ; SGP 87-17; 8 Apr 1987. CAS 65303; (3, 55–63) ; n. of Badilu; SGP 87-27; 18 May 1987. CAS 65304; (1, 42) ; Tab (Pig) I.; SGP 87-21; 15 May 1987. USNM 213161 ; (5, 23–44) ; Madang Harbor; 26 May 1970; BBC-1488 and 1490; USNM 213164 ; (4, 47–58) ; Madang Harbor; 31 May 1970; BBC-1497. Australia: Coral Sea : ANSP 123307 ; (2, 27–29) ; Big Hope Island ; TSA-33; 27 Jan 1969. ANSP 123325 ; (9, 25–30) Big Hope Island ; TSA-22; 19 Jan 1969. ANSP 123415 ; (2, 10–18) ; Little Hope Island ; TSA-1; 3 Jan 1969. ANSP 123373 ; (1, 25) ; Northern Escape Reef ; TSA-30; 24 Jan 1969. ANSP 122943 ; (3, 15–29) ; W of Little Hope Island ; TSA-32; 26 Jan 1969. ANSP 174722 ; (2, 24–26) ; Big Hope Island ; TSA-34; 27 Jan 1969. ANSP 123347 ; (2, 22–25) ; Endeavour Reef; TSA-6; 6 Jan 1969. ANSP 123316 ; (3, 38–67) ; Big Hope Island ; TSA-21; 18 Jan 1969. ANSP 123401 ; (5, 22–25) ; TSA-5; 6 Jan 1969. ANSP 122215 ; (1, 65) ; Palm Group, Curacoa Is.; 28 Sep 1952. ANSP 135333 ; (3, 56–68) ; Yonge Reef; LZ-75-56; 23 Nov 1975. CAS 40118; (2, 23–60) ; 14° 35' N 145° 36' E, Yonge Reef; 23 Nov.1975. CAS 28509; (4, 28– 37) ; 14° 0.78' S. 143° 59.0' E, Wharton Reef; 9 Oct 1960; SIO61-132 . USNM 218082 ; (3, 66–71) ; Yonge Reef; 23 Nov. 1975. CAS 40118; (2, 23–59) ; Young Reef , 14°35'S 145°36'E; 23 Nov 1975; 7– 15m. Solomon Islands: USNM 213165 ; (6, 31–54) ; 10 Mar 1965; Te Vega Cr. 6, Sta. 245. USNM 357421 ; (31, 52–77) ; Santa Cruz Is. , 10°16'30"S 166°16'30"E; 18 Sep 1998; 0– 13 m. Loyalty Islands: USNM 324847 ; (1, 70) ; Ouvea Atoll; 17 Oct. 1991. New Caledonia: USNM 324845 ; (2, 43–43) ; Noumea, Pointe Cluxel; 7 Nov. 1991. Tonga Islands : USNM 341604 ; (7, 37–54) ; JTW 93-22; 5 Nov. 1993. USNM 341605 ; (4, 45–61) ; JTW 93-17; 3 Nov. 1993. USNM 341606 ; (5, 52–70) ; JTW 93-29; 11 Nov. 1993. USNM 341607 ; (1, 44) ; JTW 93- 42; 17 Nov. 1993. USNM 341608 ; (4, 33–36) ; JTW 93-11; 28 Oct. 1993. USNM 341609 ; (1, 35) ; JTW 93-15; 2Nov. 1993. Tongatapu Group : USNM 334704 ; (2, 51–64) ; N. Of Atata I.; 28 Oct 1993. USNM 334705 ; (5, 65–73) ; Malinoa I.; 27 Oct 1993. USNM 341628 ; (6, 40–78) ; JTW 93-8; 27 Oct. 1993. Fiji: CAS 83888; GVF 2008 ; (1, 60) S. Yasarwas Is.; 4 Aug 1959. CAS 224788 ; (1, 38) ; Muaivuso , 18°08.836'S, 178°21.988'E; G02-52; 11 Feb 2002; 11– 15 m. CAS 224789 ; (5, 37–54) ; S of Kia I., 16°16.911'S, 179°09.335'E; G02-121; 30 Mar 2002; 6 m. CAS 223757 ; (2, 53–59) ; E of Nananu Passage , 17°15.063'S, 178°13.445'E; G02-82; 16 Mar 2002; 1– 5 m. USNM 242056 ; (1, 49) ; Viwa I.; VGS 82-31; 27 May 1982. USNM 262432 ; (7, 41–70) ; Naviti I.; VGS 82-32; 28 May 1982. USNM 288131 ; (3, 20–48) ; Rotuma; VGS 86-9; 14 May 1986. USNM 288195 ; (7, 28–65) ; Rotuma; VGS 86-6; 12 May 1986. USNM 262617 ; (22, 25–68) ; Lua Is., Yangasm Cluster; VGS 82-16; 2 May 1982. USNM 262455 ; (29, 39–73) ; Totoya I.;VGS 82-9 27 April 1982. Samoan Islands: American Samoa: BPBM 27929 ; (6, 62–75) ; Tutuila; 0.5– 6m.; 2 Mar. 1972. USNM 58522 ; (1, 60.1) Tutuila; cleared & stained. Kiribati: Onotoa: USNM 167318 ; (9, 31–60) . USNM 167319 ; (3, 36–63) . BPBM 15207 ; (3, 40–57) ; 1951. Society Islands: Bora-Bora : CAS 83882; GVF 1364 ; (1, 56) ; 10 Aug 1957; Sta. 32. CAS 83883; GVF 1367 ; (2, 29–46) ; 14 Aug 1957; Sta. 35. CAS 83876; GVF 1366 ; (1, 53) ; 12 Aug 1957; Sta. 34. CAS 83886 GVF 1369 ; (2, 51–55) ; 16 Aug 1957; Sta. 37. SU 24737 ; (1, 59) ; 26 Feb 1929; Huahine : CAS 83972; GVF 1358 ; (6, 34–44) ; 2 Aug 1957; Sta. 26. CAS 83889; GVF 1357 ; (1, 42) ; 2 Aug 1957; Sta. 25. Moorea : BPBM 6166 ; (26, 35–69) ; 1.5–2.5m.; 15 Sep. 1967. CAS 83807; GVF 1308 ; (13, 42– 63) ; 30 Apr 1957; Sta. 232. Raiatea : ANSP 85881 ; (4, 22–53) . CAS 83897; GVF 1359 ; (2, 48–58) ; 3 Aug 1957; Sta. 27. Tahiti : BPBM 6101 ; (1, 66) ; Papara; color, black and white; 22 m; 4 Sep 1967. CAS 83806; GVF 1339 ; (32, 23–58) ; 1July 1957; Sta. 7. CAS 83898; GVF 1344 ; (4, 51–60) ; 8 July 1957; Sta. 12. CAS 83801; GVF 1342 ; (50, 28–63) ; 4 July 1957; Sta. 10. CAS 83884; GVF 1346 ; (1, 67) ; 9 July 1957; Sta. 14. CAS 83887; GVF 1340 ; (1, 43) ; 2 July 1957; Sta. 8. CAS 83802; GVF 1347 ; (42, 37–61) ; 10 July; Sta. 15. SU 24586 ; (2, 28–59) ; 18 Feb 1929. Tuamotu Archipelago: Raroia : BPBM 9313 ; (1, 29) ; Cruise 54, Charles E. Gilbert; 1 Nov 1961. CAS 83812; GVF 99 ; (1, 63) ; 4–8 Aug 1952; Sta. 45. CAS 83962; GVF 103 ; (3, 62–72) ; 8 Aug 1952; Sta. 49. Mangareva : USNM 65426 ; (17, 68–80) ; 1 Feb 1904 -05; Albatross ; BPBM 14283 ; (6, 65–76) ; 11 Dec 1970. Line Islands: BPBM 7791 ; (13, 34–48) ; Fanning I.; 29 Oct 1968. BPBM 14254 ; (3, 28–46) ; Kiritimati; 0– 2m.; 24 Mar. 1971. CAS 83967; GVF 47 ; (1, 65) ; Palmyra I; 18 Aug 1951; Sta. 47. CAS 83971; GVF 46 ; (3, 48–68) ; Palmyra I; 17 Aug 1951; Sta. 46. Rapa : BPBM 12844 ; (1, 78) ; 28 Jan 1971. BPBM 17318 ; (2, 20–53) ; 14 Feb 1971.

Diagnosis. A Nectamia lacking dark saddles on body, caudal peduncle with a dark, wide bar, mostly above the lateral line, caudal fin with dark margins, cheek mark, wide and triangular(Tab. 2); 13 pectoral-fin rays; total gill rakers usually 26–30 ( Tab. 8); body depth 39–44%, caudal-peduncle depth 16–20%, second anal-spine length 17–21%, and pectoral-fin length 26–29% of standard length (Tab. 3).

Description. Range of proportions (as percentages of standard length) in Table 3 with paratypes and nontype material in parentheses.

Dorsal fin VII(I)–I,9; anal fin II,8; pectoral fin usually 13-13 rarely 13-14; pelvic fin I,5; principal caudal rays 9+ 8; pored lateral-line scales 24; transverse scale rows above lateral line 2; transverse scale rows below lateral line 6; median predorsal scales 3, rarely 2; circumpeduncular scale rows 12 as 5+2+5; total gill rakers 29 (24–31), usually 27 (25–30) well developed, upper arch 2+5 (0-3+5-8), lower arch 22+0 (18-22+0-1).

Villiform teeth in several rows on the premaxilla; two rows on the dentary; one row on the palatine and vomer; none on ectopterygoid, endopterygoid or basihyal.

Vertebrae 10+14. Five free hypurals, one pair of slender uroneurals, three epurals, a free parhypural. Three supraneurals, two supernumerary spines on first dorsal pterygiophore. Basisphenoid present. Supramaxilla absent. Posttemporal serrate on posterior margin. Preopercle serrate on vertical and horizontal margins. Infraorbital shelf present on third bone.

Scales ctenoid on cheek, subopercle, opercle, nape, isthmus, base of pelvic fin with two large weakly ctenoid scales, no axillary scale, ctenoid scales on base and behind pectoral fin. Pored lateral-line scales complete, fourth scale with two pores above and one pore below main canal.

*= holotype

Life colors. See Figure 6F . Head, body and fins without stripes; head with broad dark oblique cheek mark from eye to angle of preopercle, lachrymal and anterior infraorbital with yellowish mark, iris brownish with white inner ring; head and body a uniform bronze to brownish background becoming silvery below eye and on abdomen, sometimes with faint pale bars on side or a saddle under the second dorsal fin; caudal peduncle with a dark basicaudal mark mostly above pored lateral-line scales, absent below pored lateral-line scales, partial pale region just anterior to peduncle mark; first dorsal fin with dusky distal membranes between spines II–IV, second dorsal with dusky membranes between the spine and second ray, anal fin with whitish second spine and first anal ray, pectoral and pelvic rays pale, edge of caudal fin whitish with blackish interior line, rest of fin pale.

Preserved color pattern. Adults: triangular subocular cheek mark behind upper jaw; first dorsal fin with dark membrane between spines 1–4; edges of caudal-fin lobes dark; caudal peduncle with an incomplete dark, wide bar mostly at or above the lateral line, no dark saddles on body (Pacific material occasionally with a saddle under the second dorsal fin); body without pale bars or with faint pale bars on side; stomach and intestine black; roof of mouth with scattered tiny melanophores not extending onto gill arches. Juveniles: similar to the adults.

Distribution. Known from the Indian Ocean (absent in the Red Sea) to the Central Pacific ( Fig. 13 ).

Remarks. Kuiter and Kozawa (1999) provide a number of underwater photographs (p. 33, A–E) identified as Apogon savayensis . Their Apogon species 18 (p34, A–B) may be this species. Günther (1872) clearly states that there are “...no bands on the body”. No markings were detected on the body from independent examinations of E. A. Lachner and T. H. Fraser of the type. Some specimens from the West Pacific have a tendency to have a saddle under the second dorsal fin. Average gill-raker counts vary among geographic regions ( Fig. 13 , Tab. 8) by 1–1.5 elements. No obvious preserved color pattern differences were observed. This species has been confused with Nectamia luxuria which overlaps in geographic distribution except for the western Indian Ocean ( Fig. 7 ). Nectamia savayensis has a wider cheek mark and almost always lacks pale bars (when present, faint and few) in contrast to the color pattern of N. luxuria . Lachner (1951, Plate 19a–c) provided photographs of Nectamia savayensis , but his description included 9 lots of N. luxuria .

Nectamia savayensis has been collected at the stations with N. fusca (ANSP 174727, 174728), N. luxuria (MCZ 38496, CAS 83800, USNM 341618, 341622, 341626) and N. viria (USNM 263249, 268256, 341613, 341619, 341627). Five specimens in USNM 213113 have buccal enlargement, one with eggs (65 mm SL) and 4 others (64–73 mm SL) without eggs in the mouth. One 57 mm SL specimen in USNM 213187 and 2 specimens in CAS 35369 have buccal enlargement but no eggs were found. The largest of the 1963 specimens was 92 mm SL.

Figures

FIGURE 6. Underwater photographs of eight species of Nectamia. A. N. similis, new species, Calamian Is., Philippines, G. R. Allen. B. N. luxuria, new species, Andaman Sea, Thailand, U. Satapoomin. C. N. viria, new species, Raj Ampat, Indonesia, R. F. Myers. D. N. zebrinus, Red Sea, Egypt. R. F. Myers. E. N. annularis, Red Sea, Sudan, J. E. Randall. F. N. savayensis, Solomon Sea, Bougainville I., J. E. Randall G. N. bandanensis, Ogasawara Is., Japan, J. E. Randall. H. N. fusca, Red Sea, R. H. Kuiter.

FIGURE 7. Distribution of the material examined for Nectamia annularis and N. viria with regional total gill raker averages.

FIGURE 13. The distribution of collection sites for Nectamia zebrinus () N. savayensis with regional total gill raker averages.

FIGURE 18. A hypothesis for phylogenetic relationships within Nectamia based characteristics in Tables 12 and 13. One of six best trees retained based on a heuristic search using parsimony was identical to the 50% majority rule consensus of 6664 trees. All 11 characters of equal weight were parsimony-informative. Holapogon maximus was used as an outgroup.