Nebria (Falcinebria) chugokuensis, Sasakawa, 2020

Sasakawa, Koji, 2020, Taxonomic studies of the ground beetle subgenus Falcinebria Ledoux & Roux, 2005 (Coleoptera, Carabidae, Nebria) from Honshu, Japan, ZooKeys 902, pp. 37-60 : 37

Nebria (Falcinebria) chugokuensis sp. nov. Figs 16 , 34 , 35 Japanese name: Chûgoku-hime-marukubi-gomimushi

Nebria reflexa : Uéno (1985): 56 (part); Nakane (1963b): 19 (part).

Nebria (Falcinebria) reflexa uenoi : Yoshiake et al. (2011): 33 (part).

Notes.

This species is widely distributed in the Chûgoku Mountains. It is readily distinguished from other species previously considered N. reflexa distributed in Honshû by the concavity of the ventral surface of the aedeagal apex. It is distinguished from N. hikosana , described from nearby northern Kyûshû, by two pairs of setae on the ventral side of the sterna, versus one pair in N. hikosana .

Description.

Body length: ♂, 9.97-10.32 mm (mean ± SD: 10.12 ± 0.13 mm, n = 5); ♀, 10.69-11.75 mm (mean ± SD: 11.22 ± 0.53 mm, n = 3). PPW/EL: ♂, 0.312-0.324 (mean ± SD: 0.320 ± 0.005, n = 5); ♀, 0.305-0.317 (mean ± SD: 0.313 ± 0.007, n = 3). Ventral surface of aedeagal apex deeply concave. Dorsobasal lobe absent. Dorsomedian lobe absent. Dorsoapical lobe with the basal part protruding dorsally; the protrusion as long as and narrower than the right laterobasal lobe; apical portion directed anteroventrally. Right laterobasal lobe large, with the width from a ventral view wider than the width of the gonopore protrusion from a lateral view. Left laterobasal lobe large, with the width from a ventral view wider than the width of the gonopore protrusion from a lateral view. Right lateroapical lobe bifurcated at the middle, with one apex directed posterolaterally and the other directed anterolaterally; the posterolateral apex further bifurcated. Left lateroapical lobe bifurcated at the middle, with one apex directed posterolaterally and the other directed anterolaterally; the anterolateral apex further bifurcated. Ventrobasal surface without swellings adjoining the laterobasal lobes; ventrobasal swelling large, semispherical from a lateral view.

Type materials.

Holotype: ♂ (NARO), Mount Azuma, Hiwa-chô, Shôbara-shi, Hiroshima Prefecture, Japan, 27.vii.1976, S. Nakamura leg. Paratypes (NARO): 1♂, same description as the holotype; 2♂3♀, Sandan-kyô, Akiôta-chô, Hiroshima Prefecture, Japan (1♂1♀, 30.v.1957, K. Baba leg.; 1♂1♀, 31.v.1957, K. Baba leg.; 31.v.1970, I. Okamoto leg.); 1♂, Chômon-kyô, Atou, Yamaguchi-shi, Yamaguchi Prefecture, Japan, 18.v.1975, K. Tanaka leg.

Etymology.

The specific name refers to the Chûgoku Mountains, where this new species is distributed.

Figures

Figures 5 - 16. Habitus dorsal view of Nebria spp. 5 N. reflexa, a male from the type locality 6 N. sagittata sp. nov., holotype male 7 N. iidesana sp. nov., holotype male 8 - 10 N. niohozana males from Mount Chokai (8), Doai, a locality close to the type locality designated for the lectotype (9), and Mount Hakusan (10); 11 N. furcata sp. nov., holotype male 12 N. pisciformis sp. nov., holotype male 13 N. kuragadakensis sp. nov., holotype male 14 N. dichotoma sp. nov., holotype male 15 N. uenoi, a male from the type locality 16 N. chugokuensis sp. nov., holotype male.

Figures 31 - 35. Right lateral (a), ventral (b), dorsal (c), and posterodorsal (d) views of the endophallus and left laterodorsal view of the dorsoapical lobe (e) of Nebria spp. 31 N. dichotoma sp. nov., holotype male 32 N. dichotoma sp. nov., paratype male from the type locality 33 N. uenoi, male from the type locality 34 N. chugokuensis sp. nov., holotype male 35 N. chugokuensis, paratype male from Sandan-kyo. ac, concavity of aedeagal apex; remaining abbreviations are provided in Figures 17 - 21.

Figures 17 - 21. Right lateral (a), ventral (b), dorsal (c), and posterodorsal (d) views of the endophallus and the left dorsolateral view of the dorsoapical lobe (e) of Nebria spp. 17 N. reflexa male from the type locality 18 N. sagittata sp. nov., holotype male 19 N. sagittata sp. nov., a paratype male from " Asahi-mura " 20 N. iidesana sp. nov., holotype male 21 N. iidesana sp. nov., paratype male from Mount Kitamata. Abbreviations: da, dorsoapical lobe; dm, dorsomedian lobe; go, gonopore; gp, gonopore protrusion; la, left lateroapical lobe; lb, left laterobasal lobe; ra, right lateroapical lobe; rb, right laterobasal lobe. Asterisk indicates that the gonopore protrusion or lobes are not fully everted.