Mortoniella (Mortoniella) paraenchrysa Sykora, 1999

Blahnik, Roger J. & Holzenthal, Ralph W., 2017, Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *, Insecta Mundi 2017 (602), pp. 1-251 : 30-31

Mortoniella (Mortoniella) paraenchrysa Sykora, 1999

Fig. 15

Mortoniella n. sp. 3, Flint 1996: 383 [member of enchrysa subgroup].

Mortoniella paraenchrysa Sykora 1999: 381 [member of enchrysa subgroup]; Blahnik and Holzenthal 2008: 70 [member of bilineata group]; Blahnik and Holzenthal 2011: 63 [member of enchrysa subgroup].

This species is probably most readily diagnosed by its golden color and the form of the inferior appendage, which has an elongate, narrow dorsal projection that is very strongly and almost hemispherically curved. The projection is somewhat similar to that in M. denticulata Sykora , which differs in so many other respects (the separated and angulate ventrolateral projection of segment X, spines on the apex of the dorsal phallic spine, and minutely spined membranous lobes of the endophallic membrane) that it is hardly likely to be confused.

Adult —Length of forewing: male 6.4 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0:4:4. Dorsal side of forewings, head, basal segments of antennae, and legs golden-orange; ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, and palps dark brownish-black (fuscous). Wing membrane not apparently infuscated. Tibial spurs brownish-black, contrasting with legs. Wing bars absent.

Male genitalia —Ventral process of segment VI posteriorly projecting, very short, narrow basally, length about 1½ times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X with basal part slightly inflated, distinctly set off from apical part, tergum moderately elongate, lateral margins rounded, ventrolaterally with short acute lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, emarginate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with broad U-shaped connection near apex (mesal notch distinct); tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short narrow setose dorsolateral lobes and paired ventromesal lobes, each with narrow, strongly hemispherically curved, dorsal process and short acute apicomesal process. Mesal pockets of inferior appendage with very elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendages elongate, narrow, slightly widened preapically, apices acute, subequal in length to dorsal phallic spine; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin curved basally, slightly undulate in middle, and nearly rectilinearly upturned in apical ¼, apex of spine rounded; base of spine narrow, undulately curved and stalk-like, abruptly and very strongly widened on ventral margin in basal ½, forming obtusely angular ventral projection, narrowing apically from projection; spine, as viewed dorsally, somewhat widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with elongate, lightly sclerotized, lobes, extending about same length as paramere appendage, lateral margins of lobes subparallel, apices mesally curved. Endophallic membrane with conspicuous, membranously pleated, lateral lobes; phallotremal spines absent.

Material examined — PERU: Cuzco: Paucartambo, E Buenos Aires, km 135, 13.13333° S, 71.55000° W, 2150 m, 28-29.viii.1989, N Adams– 1 male (NMNH).

Distribution — Bolivia, Peru.

Figures

Figure 15. Mortoniella (Mortoniella) paraenchrysa Sykora, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral.