Montenegrina grammica erosszoltani, Feher, Zoltan & Szekeres, Miklos, 2016

Feher, Zoltan & Szekeres, Miklos, 2016, Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae), ZooKeys 599, pp. 1-137 : 28

Taxon classification Animalia Stylommatophora Clausiliidae

Montenegrina grammica erosszoltani ssp. n. Fig. 15G

Diagnosis.

Very small subspecies with dorsolateral lunella, missing basalis and non-connected anterior plica superior.

Description.

The very small, somewhat conical, greyish-corneous shell consists of 7½ to 9 whorls. The entire surface is strongly costate, with denser ribs behind the aperture and toward the apex. The deep neck inflection extends to the lateral side. The basal and peripheral crests are well developed. The ovoid to circular aperture has broadly attached, simple margin. The lamellae superior starts at the same depth as that of the end of the spiralis, occasionally with a slight overlap. The lamella inferior and medium-bent subcolumellaris are only rarely and slightly visible in front view. The broad lunella is dorsolateral. The basalis and subclaustralis are missing, the sulcalis is present. The weak anterior part of the plica superior initiates far from the lunella complex. Only the parietal edge of the clausilium plate is visible through the aperture.

Dimensions

(in mm). Holotype Hs: 8.9, Ws: 3.0, Ha: 2.4, Wa: 2.1; paratypes (HNHM 99513, n = 12): Hs: 8.5-11.4 (mean 9.5, S.D. 0.77), Ws: 2.6-3.3 (mean 2.9, S.D. 0.21), Ha: 2.3-2.9, Wa: 1.9-2.4.

Differential diagnosis.

The new subspecies differs from Montenegrina grammica grammica by its strongly costate whorls and the presence of the anterior plica superior, whereas from Montenegrina grammica improvisa by its smaller size, costate surface, deeper ending lamella subcolumellaris, and missing basalis.

Type locality.

Albania, Mat District, W of the Shkëmb i Skanderbeut (1 km from the Peshkopi to Burrel road toward Fushë-Lurë), 1160 m, 41.6461°N, 20.1807°E.

Type material.

Type locality, leg. ZE, ZF, AH, DM, 13.iv.2006, holotype (HNHM 99512), paratypes (HNHM 99513/52, ER/52, HU/52, SZ/5); same locality, leg. ZE, ZF, 11.x.2005, paratypes (HNHM 99514/138+5aj, MMM-B01327/138, ER/138, SZ/10); same locality, leg. ZF, DM, ZU, 18.v.2010, paratypes (HNHM 99515/3a+75, NHMW 111233/10); on the Peshkopi to Burrel road, 1 km W of the Fushë-Lurë junction, 1090 m, 41.6451°N, 20.1823°E, leg. ZE, ZF, AH, DM, 13.iv.2006, paratypes (HNHM 99516/27, ER/27, HU/27, SZ/5); Mali i Dejës, limestone hill of 1700 m height above the headwaters of the Lumi i Varoshit, 1540 m, 41.6746°N, 20.2112°E, leg. ZF, DM, ZU, 18.v.2010, paratypes (HNHM 99517/3+1a, SZ/2).

Etymology.

The new subspecies is named after our friend and colleague Zoltán Erőss, who accompanied the first author on several Balkan field trips, including the one during which this taxon was discovered.

Distribution.

Southern part of the Lura-Dejë mountain group, within the catchment area of the Varosh Stream (Fig. 17).

Remarks.

At one of the localities it was found syntopically with Montenegrina skipetarica puskasi ssp. n.