Mniotype adusta poltavskyi Volynkin, Matov & Behounek

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32 : 6-7

Mniotype adusta poltavskyi Volynkin, Matov & Behounek , ssp. n.

( Figs 17, 18 , 61, 62 , 102 )

Type material. Holotype: male, 27.vi. [19]63, Armenia, Sevan district, Semyonovka vill., fields, 1500 m. Slide AV0732 Volynkin (Coll. SZMN). Paratypes. 2 male, 1 female, with the same data as the holotype (Coll. SZMN); 1 male, 1 female, [ Russia], Karachaevo-Cherkessia, KGPBZ, upper course of B. Adzhara river, Kvata pass vicinity, 2300 m, alp. meadows, at light “Petromax”, 11.vii.2006, V. Shchurov (Coll. ZISP); 1 male, [ Russia, North Osetia], Star. Lars, at light, low course of Tagaurka river, 20.v.1922, M. Rjabov (Coll. ZISP); 2 females, [ Russia, North Osetia] North Osetinsky Reserve, Tsej canyon, Buron, 14-18.viii. [19]81, A. Kondratjev, ex coll. A.V. Nekrasov (Coll. ZISP); 1 male, [ Russia, Kabardino-Balkaria], KB ASSR, Terskol, 21.vii.1971, B.A. Barsov (Coll. ZISP); 1 male, [ Russia], Daghestan, Agvali, vill. Tsumada, 1800 m, 23–27.viii. [19]81, A. Bershtein, Gen. N 416, ex coll.

A.V. Nekrasov (Coll. ZISP); 1 male, 2 females, [ Georgia, Abkhazia], Sukhumi, 2.viii.1945, 27.vi.1951, 16.vii.1955, E. Miljanovskij (Coll. ZISP); 1 male, [ Georgia], Lagodechi, 18.vi. [18]80, Mlok [Mlokosevich] (Coll. ZISP); 1 male, [ Georgia], Lagodekhi, 2400 [m], 14.vii.1959, [E. Miljanovskij] (Coll. ZISP); 1 female, [ Georgia], Borshom, 20.vii.1896, [unknown collector] Coll. Gr. Prince Nikolaj Mikhajlovich (Coll. ZISP); 1 female, [ Georgia], Tskhro-Tskharo [Mt.], 8000 ft., 23.vi. [19]16, I. Filipjev (Coll. ZISP); 1 male, 3 females, [ Armenia], Gedzhanan, forest, 8000 ft., 14–19.vii.1939, [M. Rjabov] (Coll. ZISP); 1 male, Armenia, Dzhermuk, 5.vi. [19]95, M. Kalashan, ex coll. A.V. Nekrasov (Coll. ZISP); 5 males, 5 females, [ Armenia], Mal. Kavkaz, Janshakh, Kel’bodsh, 2000 m, 5.vii.1965, Effendi, ex coll. A.A. Bundel (Coll. ZISP); 1 female, [ Azerbaijan], Kussari, [18]84, Leder (Coll. ZISP); 1 female, [ Azerbaijan, Nakhichevan], Sultanbek, 21.vii.1936 [M. Rjabov] (Coll. ZISP); 1 male, Azerbaijan, Greater Caucasus, Kakh District, Ilisu, 16.vi. [19]69. I. Degtyaryov, ex coll. A.V. Nekrasov (Coll. ZISP); 1 female, Russia, North Osetia, N Caucasus,Tsei, 15–30.vii.1989, A. Belyanov (Coll. PGM); 3 males, USSR, ChIASSR, Kezenoy-am, 2000 m, 4–6.vi.1990, Á. Uherkovich, L. Ábrahám, T. S. Korolj (Coll. PGM); 1 female, Armenia, Khosrov, 10–12.vi.2003, 1600–2500 m, M. Danilevsky (Coll. PGM); 2 females, Georgia, Mingrelia, Labarde Montes Egrinsskij, Cauc. centr., 1800–2000 m, 23–26.vii.1990, K. Gaskó (Coll. PGM).

Slides AV0732 Volynkin, 0 296 Matov (males), 0 298 Matov (female).

Other material examined: 9 females, [ Russia, Karachaevo-Cherkessia], fl. et loc. Teberda, 4200 ft., 6.vii- 9.viii.1912, S. Tschetwerikow (Coll. ZISP); 1 male, [ Russia], North Osetia, Tsejsk. ushch. [Tsej canyon], 13.viii.1931, at light, M. Rjabov (Coll. ZISP); 1 male, [ Russia, North Osetia], SO ASSR, Alagirskij district, Tsej, 7.viii.1976, A.N. Poltavskij (Coll. ZISP); 1 female, [ Georgia], Caucasus, Ritsa, 20.vii. [19]70, E. Miljanovskij (Coll. ZISP); 1 male, [ Georgia], Lagodekhi, 7.viii.1956 [E. Miljanovskij] (Coll. ZISP).

Note. Turkish specimens ( Fig. 19 ) are externally more close to M. adusta adusta than to M. adusta poltavskyi and have male genitalia ( Fig. 65 ) without notable differences from the nominotypical subspecies. The correct establishing of subspecific belonging of Turkish populations needs the examination of extensive material from various areas of Turkey.

Diagnosis. The new subspecies differs clearly from the nominotypical race by broader forewings and welldefined contrast pattern.

Description. External morphology ( Figs 17, 18 ). Wingspan 35–46 mm. Head, thorax and abdomen dark brown or blackish. Forewing relatively broad. Ground colour of forewing reddish brown in males and blackish brown in females. Pattern well-defined, contrast. Crosslines thin, double. Submarginal line pale, thin, strongly dentate. Marginal line present as a series of black dots between the veins. Basal dash short, thin, black. Claviform dark brown or blackish, with thin black border; medial fascia thin, black. Medial area shadowed around claviform and medial fascia and in the cell also. Terminal area dark brown or blackish brown. Subterminal area inwards from the submarginal line with dark brown or blackish cuneate streaks between the veins. Cilia dark brown. Hindwing paler in males, brownish grey, discal spot diffuse, semilunar. Cilia pale grey. Male genitalia ( Figs 61, 62 ): show no significant differences from M. adusta adusta . Female genitalia ( Fig. 102 ): have a somewhat broader ostium bursae and a somewhat longer and slightly more curved ductus bursae in comparison with those of the nominotypical subspecies.

Distribution. Caucasus, Transcaucasia.

Etymology. The subspecies name is dedicated to Dr. Alexander Poltavsky, a well-known expert on Caucasian Noctuidae .

Figures

FIGURES 11 – 20. Mniotype spp. adults: 11, M. lama ayubaevorum, holotype male, Russia, Altai Mts (ZISP); 12, ditto, paratype male, Russia, Altai Mts (ZISP); 13, ditto, paratype female, Russia, Altai Mts (AVB); 14, ditto, paratype female, Russia, Altai Mts (AVB); 15, M. adusta adusta, male, Germany (GBG / ZSM); 16, ditto, female, Russia, Minussinsk (ZISP); 17, M. a. poltavskyi, holotype male, Armenia (SZMN); 18, ditto, paratype female, Russia, North Osetia (ZISP); 19, M. adusta? ssp., Turkey (HMNH); 20, M. adusta? ssp., SE Kazakhstan, (GBG / ZSM).

FIGURES 57 – 62. Mniotype spp., male genitalia: 57, M. lama ayubaevorum, holotype, Russia, Altai Mts, slide AV 0457 Volynkin; 58, M. l. ayubaevorum, paratype, Russia, Altai Mts, slide AV 0504 Volynkin; 59, M. adusta adusta, Russia, Moskow reg., glyc. prep. Volynkin; 60, M. adusta adusta, Russia, Altai Mts, slide AV 0516 Volynkin; 61, M. adusta poltavskyi, holotype, Armenia, slide AV 0732 Volynkin; 62, M. adusta poltavskyi, paratype, Armenia, slide AV 0733 Volynkin.

FIGURES 97 – 104. Mniotype spp., female genitalia: 97, M. lama lama, Kyrgyzstan, slide AV 0033 Volynkin; 98, M. l. kortka, paratype, SE Kazakhstan, slide AV 1122 Volynkin; 99, M. l. etugen, paratype, W Mongolia, slide AV 0673 Volynkin; 100, M. l. ayubaevorum, paratype, Altai Mts, slide AV 0505 Volynkin; 101, M. adusta adusta, Altai Mts, slide AV 0724 Volynkin; 102, M. a. poltavskyi, paratype, N Caucasus, slide 0 296 Matov; 103, M. a. sommeri, Iceland, slide 0 294 Matov; 104, M. adjuncta adjuncta, holotype, Dalhousie, slide A. Giusti (© BMNH) (photo by A. Giusti).

FIGURES 63 – 68. Mniotype spp., male genitalia: 63, M. adusta sommeri, Greenland, slide 0 324 Matov; 64, M. a. sommeri, Iceland, slide 0 293 Matov; 65, M. adusta? ssp., Turkey, slide AV 0778 Volynkin; 66, M. adusta? ssp., SE Kazakhstan, slide 5838 Behounek; 67, M. adjuncta adjuncta, N India, Himalaya Mts, slide GB 12103 Behounek; 68, M. a. adjuncta, Pakistan, Himalaya Mts, slide AV 0792 Volynkin.