Mitella stylosa var. stylosa

M. Wakabayashi, 2006, Tiarella and Mitella, Flora of Japan, Volume 2 b: Angiospermae-Dicotyledoneae: Archichlamydeae (b), Kodansha, pp. 70-75 : 73

var. stylosa

Mitella japonica Maxim, var . integripetala Makino in Bot. Mag. Tokyo 19: 17 (1905).

Mitella stylosa H.Boissieu var . integripetala (Makino) Ohwi in Acta Phytotax. Geobot. 1: 64 (1932).

Mitella stylosa H.Boissieu f. integripetala (Makino) H.Hara in Nakai & Honda, Nov. FI. Jap. 3: 132 (1939).

Mitella leiopetala Ohwi et Okuyama in Bull. Natl. Sci. Mus. Tokyo no. 33: 112 (1953).

Japanese name: Takimi-charumeru-so.

Perennial herbs. Rhizomes somewhat thickened, creeping, with a tuft of leaves at apex. Radical leaf blade ovate, cordate, subacute or obtuse, 4-9 cm long, 3-7 cm wide, shallowly or obsoletely lobulate, irregularly toothed, upper surface sparsely hirsute, dark green, lower surface papillose-pilose, pale green often with purplish tinge; petiole 7-15 cm long, hirsute, with fused axillary stipules at base; stipules scarious, oblong, sparsely minutely hairy along margin. Flowering stems ca. 25 cm tall, erect, densely glandular pubescent, hirsute toward base. Inflorescences racemes, many-flowered. Flowers April; pedicel ca. 2 mm long, minutely glandular. Calyx tube campanulate-obconical, densely glandular pubescent, brownish; calyx lobes 5, triangular, acute, nearly glabrous adaxially, glandular abaxially, ca. 1.4 mm long, erect or slightly recurved near tip, brownish purple. Petals 5, subulate-linear or rarely with 1 (or 2) short lobes on one or both sides, glabrous, 1.5-3 mm long, often ca. 5 mm long and 3(or 4)(or 5)-pinnatiparted with obliquely ascending pinnae and glandular dots, reflexed, reddish purple. Stamens 5, alternate calyx lobes, adjacent to base of petals, 1-1.4 mm long; anthers broadly ovoid, cordate, ca. 1 mm long at anthesis, yellow. Disc inconspicuous. Ovary inferior. Styles 2, erect, thick at base, ca. 1.4 mm long; stigmas thickened, 4-lobed. Seeds oblong, ca. 1.1 mm long, scabrous and densely papillose, brownish.

Chromosome number: 2n = 28 (Wakabayashi 1973).

Japan: C. Honshu (Ibuki and Suzuka Mountains in Shiga and Gifu Pref.). Moist, shady places along streams in mountainous areas. Endemic.

Icones: Terasaki, rev. ed.: f. 1157; Wakabayashi in Acta Phytotax. Geobot. 28: 113, f. 1 (1977); Satake et al., Herb. PI. 2: photo. 152 5 & 6.