Megalothoraca ferruginea ( Enderlein, 1912 )

Wendt, Lisiane Dilli & Marinoni, Luciane, 2019, Taxonomic revision of Megalothoraca (Diptera: Richardiidae) with description of a new species, synonyms and new combination, Zoologia (e 31456) 36, pp. 1-15 : 4

Megalothoraca ferruginea ( Enderlein, 1912)

Figs 1–4

Phlebacrocyma ferruginea Enderlein, 1912: 106 . Type locality: Santa Inez, Ecuador. Hennig 1938a: 121 (discussion); Aczél 1950: 44 (catalogue).

Megalothoraca ferruginea Steyskal, 1968: 5 (catalogue).

Diagnosis and comments. Megalothoraca ferruginea , known from a female specimen, is the only species within genus with head and thorax entirely reddish brown, fore and mid coxae, fore and mid femora pale brown ( Figs 1, 3) (black in M. undulosa and most other species, except yellow in M. flava ).

Redescription. Holotype female, MZPW: Measurements (in mm): body length 8.20 (8.50 according to Enderlein 1912: 106); wing length 8.00 (8.25 according to Enderlein 1912: 106). Head ( Figs 1, 3). Extremely damaged, especially oral region and eyes, ratios not measured, and some parts not observed. Anterior ocellus inserted on posterior third of frons. Reddish brown, except ocellar triangle and vertex brown. Vertex in frontal view straight, not concave. Frons and postcranium sparsely whitish and brownish pollinose, respectively. Antenna brownish. Clypeus, palpus, gena and prementum missing. Ocellar seta slightly longer than orbital seta; inserted at same level. Thorax ( Figs 1, 3). Entirely reddish brown, with scutellum, subscutellum and mediotergite slightly darker. Scutum 1.33 times longer than wide. Postpronotal seta absent. Legs ( Fig. 1). Fore coxa, trochanter, and femur yellowish orange; mid and hind coxae, trochanters, and femora reddish brown; all tibiae brown; all tarsomeres yellowish. Fore and mid coxae without anterior spine. Femora similar in thickness. Left fore femur with two anteroventral and three posteroventral spine-like setae, and right fore femur with two antero- and posteroventral spine-like setae; setae different in size, occupying apical half of femur. Fore and mid coxae without spine-like seta. Mid and hind femora with one stout preapical anteroventral spine-like seta; stouter than those on fore femur. Hind coxa bare on posterior surface. Wing ( Fig. 1). Generally hyaline except for the following marks: cell bc yellowish; faint and diffuse brownish basal band from vein R 1 at level of costal break, passing on base of bifurcation between R 2+3 and R 4+5, and on vein bm-cu; oblique yellow area in cell r 1, from costal break reaching medial band; incomplete and narrow medial brown band, from costal margin to vein CuA 1, passing on spurious vein and on vein r-m, with irregular margins; and wider and conspicuous apical brown spot, faint in cell m, with proximal margin irregular, almost straight. Vein R 2+3 from apical third equally twice sinuous. Spurious vein faint and short, at same level of vein r-m. Vein M, distal to vein r-m, sinuous and curved towards R 4+5. Vein r-m at distal third of cell dm length. Halter with stem brownish and knob brown. Abdomen ( Figs 1, 3). Entirely dark brown, except contrasting tergite 5 yellowish orange. Tergites brown setulose, very sparsely whitish pollinose. Syntergite 1+2 with two pairs of weak preapical laterodorsal setae. Sternites 1–4 brownish, sternite 5 yellowish orange. Female terminalia ( Fig. 4). Not dissected. Oviscape reddish yellow. Tip of cerci rounded.

Male. Unknown.

Type-material examined. Holotype female, MZPW: “ Santa Inéz \(Ecuad. [ Ecuador])\ R.Haensch S.”; “ Type ” [red label]; “ Phlebacrocyma \ferrugineum\Type Enderl. ♀ [handwritten]\ Dr. Enderlein det. 1912” ( Fig. 2). The holotype is pinned, in poor condition ( Figs 1, 3): head damaged, with eyes, oral and genal regions, and postcranium ventrally missing, right anterior region of anepisternum and katepisternum with a hole; oviscape, apex of segment 8, and cerci exposed ( Fig. 4).

Distribution. Ecuador.