Mecynotarsus bullatus, Kejval & Cz, 2013

Kejval, Zbyněk, 2013, Taxonomic revision of the Australian Notoxinae (Coleoptera: Anthicidae), Acta Entomologica Musei Nationalis Pragae (suppl.) 53, pp. 1-98 : 28-31

Mecynotarsus bullatus sp. nov.

( Figs 33, 34 , 139 , 147, 148 , 172 , 191 )

Type locality. Australia, Western Australia, 13 km EN of Newman, 23°15′S 119°52′E.

Type material. HOLOTYPE: ♂, ‘(23.15S 119.52E) 13 km. E. by N. of Newman ,WA. 12.xi.70. E.B. Britton [p] // ANIC specimen [p; green label]’ ( ANIC) . PARATYPES: 1 ♂ 1 ♀, same data as holotype ( ANIC) ; 3 ♂♂ 6♀♀, ‘ Minilya River , WA. 23. 49S 114. 00E 29 Mar. 1971 Upton & Mitchell [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC, 2 spec. ZKDC) ; 1 ♂ 2 ♀♀, ‘ West Peawah R. WA 14.iv.1976 K. & E. Carnaby [p] // ANIC specimen [p; green label]’ ( ANIC) ; 2 ♀♀, same data, except: ‘AUST. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 2 ♀♀, ‘ W. Grey River WA 6 March 1983 K. & E. Carnaby at light’ ( ANIC) ; 1 ♀, ‘ Regans Ford W.A. 18 Oct. 1978 on Brand Hwy. at light K. & E. Carnaby [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♂ 1 ♀, ‘ Geraldton Oct ’ 31 W. A. [p] // Australia, Harvard Exp. Darlington [p] // [plain yellowish label]’ ( DCDC) ; 1 ♂, ‘ Qld. Greencale 70 Km SW at light 17-24 Jan. 96 A. J. Watts [p] // SAMA Database No. 25-029000 [p]’ ( SAMA) ; 1 ♀, ‘ Qld. Greenvale 70 km SW At light 1-10 Mar. 95 A. J. Watts [p] // SAMA database No. 25-028947 [p]’ ( SAMA) ; 2 ♀, same data, except database numbers: 25-028953 or 25-028967 ( SAMA) ; 1 ♀, ‘ Qld. Greenvale 70 km SW at light, 12-21 Apr 1995 A. J. Watts [p] // SAMA Database No. 25-028950 [p]’ ( SAMA) ; 1 ♀, ‘QLD. 70 km SW GREENVALE. 16-28 JAN 95 A. J. WATTS [p] // SAMA Database No. 25-029037 [p]’ ( SAMA) ; 1 ♀, ‘QLD. 70 km SW GREENVALE. 15-24 FEB 95 A. J. WATTS [p] // SAMA Database No. 25-029019 [p]’ ( SAMA) ; 2 ♂♂ 4 ♀♀, ‘ Upper Daintree R., Via Daintree, N. Qld. 27.xii.1964. G. Monteith’ ( QMBA) ; 2 ♂♂ 1 ♀, ‘W Normanby R., N. Qld., 40 ml. W. of Cooktown 2.i.1946. G. Monteith’ ( QMBA) ; 1 ♀, ‘ Normanton Q 20.v.1976, K. & E. Carnaby’ ( ANIC) ; 3 ♀♀, ‘ AUSTRALIA, N. Territory West MacDonnell Range Nat. Park , SIMPSON GAP 23°40′S 133°43′E, 600 m, 3.- 5.01.2009, St. Jakl leg. [p]’ ( ZKDC, NMPC) ; 1 ♂, same data, except: 11.01.2009 ( ZKDC) .

Description (holotype, male). Body length 3.3 mm. Body reddish, pronotal disc darker, reddish brown, elytra unicoloured; legs and antennae reddish.

Antero-lateral margins of frons moderately raised / lobed near antennal insertion. Gular rugules of different sizes, anteriorly larger, ordered and fused ( Fig. 139 ). Clypeal granules minute. Setation of head vertex fine, appressed, with numerous, very long, more raised setae medially, distinctly coarser, subdecumbent ventro-laterally near eyes. Antennae rather long; antennomere I long and robust, strongly enlarged apically; antennomeres III–V about twice, X 1.3 times as long as wide; setation mostly fine, distinctly coarser on basal antennomeres, especially antennomere I conspicuously setose, with coarsely fringed apical margin and numerous long bristly setae.

Pronotum moderately transverse, 1.4 times as long as wide, its lateral margins rather strongly and evenly convex in dorsal view; posterior collar very narrow and inconspicuous. Pronotal horn robust and wide, clearly triangular, its posterior angles conspicuously projecting in dorsal view ( Fig. 147 ); horn margins armed with 3 lobules on each side, posterior lobules very wide, apical lobule simple, widely rounded; horn crest very conspicuous, strongly raised, short and situated rather posteriorly, with rugules mostly fused and forming complete semicircular rim (single separate rugule posteriorly on each side); submarginal rugules mostly very coarse, distinctly spaced; single large, median longitudinal rugule and several minute granules scattered posteriorly. Setation scaly, whitish laterally, pale reddish dorsally; scales on pronotal disc of two sizes, shorter and appressed or longer and subdecumbent, both elongate, truncate apically and somewhat glossy; antebasal paired setae present medially, difficult to recognize laterally owing to presence of numerous long tactile setae.

Elytra 1.7 times as long as wide; omoplates and postbasal impression absent. Setation mixed whitish and reddish, and brownish, forming vague brownish marking ( Fig. 191 ), scaly, appressed and evenly ordered; scales linear, rounded to subtruncate apically, slightly glossy and distinctly spaced (surface clearly visible, Fig. 172 ); erect tactile setae present, more numerous and longer in basal third, especially at base and humeri, shorter and very sparsely scattered mesally, and absent in apical third.

Male characters. Sternum VII nearly simple. Tergum VIII and aedeagus as in Figs 33, 34 .

Variation. Body length (♂ ♀) 2.7–3.7 mm; some specimens with nearly unicolorous, whitish to pale reddish scales on elytra, brownish marking may be both absent and conspicuous; pronotal horn in the specimens from Queensland mostly subtriangular, with less projecting posterior angles ( Fig. 148 ).

Differential diagnosis. Mecynotarsus bullatus sp. nov. is undoubtedly close to M. hirtipennis sp. nov. as suggested by similarity of both external (triangular pronotal horn, semicircular crest rim, numerous tactile setae), and male characters. Its specimens are usually larger and more robust than those of the latter species, and differ especially by the longer, denser and somewhat double setation of the pronotal disc, with longer scales rather subdecumbent and somewhat overlapping shorter appressed scales, by the more robust antennae (basal antennomeres), and by the evenly narrowing, arcuately divergent and apically pointed parameres.

Etymology. The Latin adjective, bullatus , - a, -um (= bubble-like); named in reference to the peculiar rounded shape of horn crest rim.

Distribution. Australia: Queensland, Northern Territory, Western Australia.

Figures

Figs 33–39. 33–34 – Mecynotarsus bullatus sp. nov.: 33 – tergum VIII and apex of spiculum, 34 – aedeagus in dorsal view. 35–37 – M. canthariphilus sp. nov.: 35 – sternum VII, 36 – tergum VIII, 37 – aedeagus in dorsal view. 38–39 – M. centralis sp. nov.: 38 – tergum VIII, 39 – aedeagus in dorsal view. Scale (0.5 mm): A – Figs 33, 34, 38; B – Figs 35; C – Figs 36, 39; D – Fig. 37.

Figs 133–140. 133 – Mecynotarsus setulosus sp. nov., head in lateral view (paratype, Fowlers Gap), 134 – M. excavatus sp. nov., apex of elytra in female (paratype, Aoue-Enga), 135 – M. grandior sp. nov., apex of elytron in male (paratype, Kimberley Research Station), 136 – M. obesus sp. nov., metatibia with tarsus (paratype, Hull River), 137 – M. excavatus sp. nov., male protibia (paratype, Aoue-Enga). 138–140 – head in ventral view: 138 – M. auripilosus sp. nov. (paratype, Kambah Pool), 139 – M. bullatus sp. nov. (paratype, Regans Ford), 140 – M. ziczac King, 1869 (Swain Reefs).

Figs 141–149. Pronotal horn in dorsal view: 141 – Leptoprion angulatus Krekich-Strassoldo, 1914, 142 – Mecynotarsus albellus Pascoe, 1866 (Wemen), 143 – M. amabilis Lea, 1895 (Shoalhaven River), 144 – M. apicipennis Lea, 1922 (syntype), 145 – M. auripilosus sp. nov., 146 – M. bidens sp. nov., 147 – M. bullatus sp. nov., 148 – the same (paratype, Daintree), 149 – M. excavatus sp. nov.

Figs 168–176. Setation of elytra: 168 – Mecynotarsus amabilis Lea, 1895 (Shoalhaven River), 169 – M. apicipennis Lea, 1922, 170 – M. auripilosus sp. nov. (paratype, Kambah Pool), 171 – M. bidens sp. nov., 172 – M. bullatus sp. nov. (paratype, Minilya River), 173 – M. concolor King, 1869 (Olary), 174 – M. excavatus sp. nov., 175 – M. granulatus sp. nov., 176 – M. hirtipennis sp. nov.

Figs 186–193. Habitus: 186 – Leptoprion angulatus Krekich-Strassoldo, 1914, 187 – Mecynotarsus amabilis Lea, 1895 (Shoalhaven River), 188 – M. apicipennis Lea, 1922, 189 – M. auceps sp. nov., 190 – M. auripilosus sp. nov., 191 – M. bullatus sp. nov., 192 – M. concolor King, 1869 (Olary), 193 – M. excavatus sp. nov.