Madagotricha ranomafanae, Jaschhof & Jaschhof, 2007

Jaschhof, Mathias & Jaschhof, Catrin, 2007, Madagotricha ranomafanae gen. et sp. n., first member of the Heterotricha group (Diptera: Sciaroidea) from Madagascar, African Invertebrates 48 (2), pp. 93-100 : 96-99

Madagotricha ranomafanae sp. n.

Figs 1–4

Etymology: From the type locality, Ranomafana.

Description:

Male.

Body size 2 mm.

Head ( Figs 1A, B ): Basal antennal flagellomeres as in Fig. 1B . Labrum short subtriangular, non-setose. Maxilla with lacinia not traceable. Maxillary palpus with 2 segments; first segment smooth, bearing 1 seta; second segment slightly larger than first, beet-shaped, bearing setae and on mesial surface short hyaline sensilla.

Thorax ( Fig. 2 ): As for genus.

Wing ( Fig. 1C ): In two specimens seen, sc–r traceable as very pale vein, otherwise sc–r absent. Membrane rather sparsely covered with setae. R1 and R5 each apically with ventral setae; stem vein, Sc, R, R1 , R5 , M1 , M2 , CuA1, and CuA2 with dorsal setae .

Terminalia: Sternite 9 large, subrectangular, setose ( Fig. 3A ). Tergite 9 setose, apical lobes ventrally each with 5–7 one-pointed setae ( Fig. 3C ). Gonocoxites apicolaterally with slight projections; ventral and lateral surfaces setose except a large membranous portion ventrobasally ( Fig. 3A ). Gonostylus curved, tapered in distal third, apically rounded and bearing a few setulae ( Fig. 3B ). Tegmen ventromedially separated throughout, broadly rounded on either side of a medial cleft; apically with strong, rounded lobe extending basally into narrow process ( Figs 4B, C ). Aedeagus ventro-apically with hood-like structure and latero-apically with chamber-like structure ( Figs 4B, C ). Female. Unknown.

Holotype: ơ MADAGASCAR: Fianarantsoa Prov.: Parc National Ranomafana , radio tower, 21 ° 15'05"S: 47 ° 24'43"E, alt. 1150 m, at edge of montane rainforest, 1–19.ix.2002, Malaise trap, R. Harin’Hala Hasinjaka & M.E. Irwin ( CASC).

Paratypes: 7ơ same data as holotype ( CASC, ZIMG) .

Figures

Fig. 1. Madagotricha ranomafanae sp. n., male: (A) paratype, head, frontal view; (B) paratype, antennal flagellomeres 1–4, lateral view; (C) holotype, wing, dorsal view. Scale bars = 0.1 mm (A), 0.05 mm (B) and 1.0 mm (C).

Fig. 2. Madagotricha ranomafanae sp. n., male holotype, thorax, lateral view. Scale bar = 0.2 mm.

Fig. 3. Madagotricha ranomafanae sp. n., male paratypes: (A) terminalia, ventral view; (B) gonostylus, mesial view; (C) tergite 9, cerci and hypoproct, ventral view. Designations: 1 – sternite 9, 2 – tergite 9, 3 – gonocoxa, 4 – anterior portion of gonocoxal apodeme, 5 – gonostylus, 6 – parameral apodeme, 7 – medio-apical lobe of tegmen, 8 – aedeagus, 9 – cercus, 10 – hypoproct. Scale bars = 0.05 mm (A, C) and 0.03 mm (B).

Fig. 4. Madagotricha ranomafanae sp. n., male paratypes: (A) terminalia, lateral view; (B) tegmen and aedeagus, ventrolateral view; (C) tegmen and aedeagus, ventral view. Designations: 1 – sternite 9, 2 – tergite 9, 3 – gonocoxa, 4 – anterior portion of gonocoxal apodeme, 5 – gonostylus, 6 – parameral apodeme, 7 – medio-apical lobe of tegmen, 8 – tegmen process, 9 – dorsobasal margin of tegmen, 10 – aedeagal apodeme, 11 – chamber-like structure, 12 – hood-like structure, 13 – cercus. Scale bars = 0.05 mm.