Macroderes namakwanus Frolov & Scholtz, 2005

Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. & Sole, Catherine L., 2018, An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa, Zootaxa 4504 (1), pp. 41-75 : 63

Macroderes namakwanus Frolov & Scholtz, 2005

( Figs. 13 , 35 , 42 , 61 , 75 )

Macroderes namakwanus Frolov & Scholtz 2005: 388 .

Type locality. RSA, Western Cape Prov, Hoekbaai , 31°09'26.07''S 017°45'55.70''E .

Type material examined. Holotype, ♂ ( TMSA): “ RSA / Western Cape Prov / Hoekbaai / A. Frolov & C. Deschodt leg [printed] // AF-0039 (19) / 4–11. IX. 2003 / S 31° 04' 26.07'' E 017° 45' 55.70'' [printed] // HOLOTYPUS / Macroderes namakwanus / Frolov & Scholtz / 2003 [red label, printed]” ; Paratypes: 1♂ ( TMSA): “S. Afr / Namaquland / Brakriver mouth / 36°.06' S–17°.44' E [printed] // 25. 8. 1979 // E– Y: 1597 / groundtraps / 63 days / Endrödy—Younga leg [printed] // PARTYPUS / Macroderes namakwanus / Frolvo & Scholtz / 2003 [red label, printed]” ; 1 ♂ ( TMSA): “S. AFR / Namaquland / Hoekbaai / 2 Km / E NE / 31°. 11' S 17°. 47' E [printed] // 27. 8. 1979 // E–Y: 1612 / white sand / night / Endrödy—Younga leg [printed] // PARATYPUS / Macroderes namakwanus / Frolov & Scholtz / 2003 [red label, printed]” ; 1 ♂ ( TMSA): “S. Afr / Namaquland / Kotzerus / 30° 57' S– 17°.50' E [printed] // 23. 8. 1979 // E–Y: 1581 / white dune / day /. Endrödy—Younga leg [printed] // PARATYPUS / Macroderes namakwanus / Frolov & Scholtz / 2003 [red– printed]” ; 1 ♀ ( TMSA): “S. Afr / Richtersveld / Klein Helskloof / 28°.51' S–17°.24' E [printed] // 8.9.1976 // E–Y: 1239 / groundtraps / 32 days / Endrödy—Younga leg [printed] // groundtrap with faeces bait [printed] // PARATYPUS / Macroderes namakwanus / Frolov & Scholtz / 2003 [red label, printed]” .

Additional material examined. 2 ♂♂ and 3 ♀♀ ( UPSA): “ South Africa / 6 km N of / Kotzerus / S 30° 69' 26.11'' E 017° 86' 9.83'' / 12. 8. 2015 ” .

Size range. Males length: 10.4–12.8 mm, width: 7.0– 8.7 mm; females length: 11.7–12.5 mm, width: 8.0–9.0 mm.

Differential diagnosis. M. namakwanus is in habitus similar to M. endroedyi ( Fig. 13 ) however, it is differentiated by the combination of the following characteristics: Pronotum regularly punctate lacking the distinct tubercle in between ( Fig. 61 ) and no shiny tubercles on the elytra ( Fig. 75 )

Habitat and distribution. See distribution under M. mutilans ( Fig. 2)

Figures

FIGURES 11–14. Habitus of Macroderes species. 11, M. minutus Frolov & Scholtz, 2005; 12, M. mutilans Kolbe, 1908; 13, M. namakwanus Frolov & Scholtz, 2005; 14, M. nitidus Harold, 1877.

FIGURES 24–39. Internal sac of aedeagus (24), aedeagus (25) and internal sac sclerite (26-39) of Macroderes species. 26, M. endroedyi Frolov & Scholtz, 2005; 27, M. amplior Frolov & Scholtz, 2005; 28, M. bias (Olivier, 1789); 29, M. cederbergensis Abdalla & Deschodt, new species; 30, M. cornutus Frolov & Scholtz, 2005; 31, M. foveatus Frolov & Scholtz, 2005; 32, M. gifboomi Abdalla & Scholtz, new species; 33, M. leipoldti Abdalla & Deschodt, new species; 34, M. mutilans Kolbe, 1908; 35, M. namakwanus Frolov & Scholtz, 2005; 36, M. oreatus Abdalla & Deschodt, new species; 37, M. porselinus Abdalla, new species; 38, M. soleiana Abdalla & Deschodt, new species; 39, M. tourtosus Abdalla & Scholtz, new species.

FIGURES 40–45. Pronotum of Macroderes species. 40, M. foveatus Frolov & Scholtz, 2005; 41, M. cornutus Frolov & Scholtz, 2005; 42, M. namakwanus Frolov & Scholtz, 2005; 43, M. politulus Preudhomme de Borre, 1880; 44, M. greeni (Kirby, 1818); 45, M. endroedyi Frolov & Scholtz, 2005.

FIGURES 58–63. Punctuation of the base of pronotum of Macroderes species. 58, M. greeni (Kirby, 1818); 59, M. arrowi Janssens, 1939; 60, M. bias (Olivier, 1789); 61, M. namakwanus Frolov & Scholtz, 2005; 62, M. endroedyi Frolov & Scholtz, 2005; 63, M. politulus Preudhomme de Borre, 1880.

FIGURES 70–75. Sculpture of second elytra intervals of Macroderes species. 70, M. arrowi Janssens, 1939; 71, M. cornutus Frolov & Scholtz, 2005; 72, M. endroedyi Frolov & Scholtz, 2005;73, M. foveatus Frolov & Scholtz, 2005; 74, M. mutilans Kolbe, 1908; 75, M. namakwanus Frolov & Scholtz, 2005.