Macohasa tetraspila (Černý, 2009) Dubatolov & Volynkin & Kishida, 2018

Dubatolov, Vladimir V., Volynkin, Anton V. & Kishida, Yasunori, 2018, Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae), Zootaxa 4407 (3), pp. 383-400 : 390-391

Macohasa tetraspila (Černý, 2009) , comb. nov.

( Figs. 13–14 , 28 , 39 )

Macotasa tetraspila Černý, 2009 , Moths of Thailand, 6: 139, pl. 28, figs. 278a, 278b (Type locality: “SE Thailand, Chantaburi, Kaeng Hang MaeW Dist., 10°4’41” N, 101°48’5” E ”).

Type material examined. Holotype: ♂ ( Figs. 13 , 28 ), SE Thailand, Chantaburi, Kaeng Hang Maew Dist. , 10°4’41” N, 101°48’5” E, 2-3.VI.2005, leg. K. Černý, slide MWM 33812 Volynkin (Coll. MWM/ ZSM) . Paratypes: 7 ♂, 10 ♀, same data as in the holotype, slides AV4183 ♂, AV4184 ♀ Volynkin (Coll. CKC) ; 3 ♂, SE Thailand, Chantaburi, 265 m, Bar Ta Moon, Soi Dao distr. , 13°3’40’’N, 102°14’45’’E, 21.XI.2005, leg. K. Černý, slide AV4184 ♂ Volynkin (Coll. CKC).

Redescription. Male ( Fig. 13 ). Forewing length 9–10 mm. Forewing ground color yellowish brown, with costa slightly conVeX in subapical third and dorsum noticeably conVeX in basal half; costal margin with one broad, diffuse antemedial black spot; central cell with a long and Very narrow fold with androconial scales; posterior section of forewing posteriorly the costal spot with broad diffuse crossline presented as blackish suffusion. Hindwings ochreous. Male genitalia ( Fig. 28 ). Uncus simple, long, basally broad, apically pointed, curVed downwards at apical half; scaphium broad, heaVily sclerotized; Vinculum elongated, more or less V-like, with tapered apeX; apeX of ValVe rounded; harpe-like fold small, pimple-like; distal saccular process short, broad, with apeX pointed and curVed dorsally. Aedeagus long, slightly curVed; Vesica broad, with two membranous subbasal diVerticula, one large globular distal diVerticulum with granulation, and one long and basally curVed distal diVerticulum with granulation; Vesica ejaculatorius broad, situated laterally. Female ( Fig. 14 ). Forewing length 11– 12 mm. Forewing ground color pale yellow, with a row of 5 black spots, the middle one Very small, dot-like, sometimes absent; hindwing pale yellow. Female genitalia ( Fig. 39 ). OVipositor short, broad; apophyses long and thin, apophyses anteriores slightly longer than apophyses posteriores; ductus bursae short, narrow, membranous, with large, Ventral plate; corpus bursae broad, sack-like, membranous, with one elliptical signum; appendiX bursae short, bulbous, situated latero-posteriorly.

Remark. M. tetraspila is a closest relatiVe of M. dimorpha . Both species haVe female genitalia differing significantly from those of two other congeners by the large Ventral plate of ductus bursae and elliptical signum (whereas in M. cernyi sp. nov. and M. orientalis the Ventral plates are round and bulbous, and the signa are barshaped). The male genitalia of both seXes are similar to those of M. cernyi sp. nov. and M. orientalis , but the scaphium is significantly broader, the harpe-like fold is significantly smaller, the distal saccular process is smaller, the Vinculum is significantly larger, and both distal diVerticula of aedeagus Vesica are granulated. Differences of M. tetraspila from M. dimorpha are listed in the diagnosis of M. dimorpha .

Distribution. SE Thailand, Malaysia (Malay Peninsula) (Bucsek 2014).

Figures

FIGURES 11–19. Macohasa and Chinasa spp.: adults. 11, M. orientalis, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12, M. orientalis, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13, M. tetraspila, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14, M. tetraspila, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15, M. dimorpha, syntype ♂, Philippines, Luzon (©NHMUK); 16, M. dimorpha, syntype ♀, Philippines, Luzon (©NHMUK); 17, Ch. costalis costalis, holotype ♂, N China (©NHMUK); 18, Ch. costalis costalis, ♂, China, Zhejiang (MWM/ZSM); 19, Ch. costalis costalis, ♀, China, Guangdong (SZMN).

FIGURES 27–29. Macohasa spp.: male genitalia (genital capsules in ventral vieW). 27, M. orientalis, Malaysia, slide MWM 31775 Volynkin; 28, M. tetraspila, holotype, Thailand, Chantaburi prov., slide MWM 33812 Volynkin; 29, M. dimorpha, Philippines, PalaWan, slide MWM 31777 Volynkin.

FIGURES 36–41. Lithosiina spp.: female genitalia (ventral vieW). 36, Prabhasa venosa, [India, Meghalaya] Khasis, slide BMNH(E) Arct-4557f HolloWay (©NHMUK); 37, Macohasa cernyi, paratype, N Vietnam, slide MWM 31774 Volynkin; 38, Macohasa orientalis, Malaysia, slide MWM 31776 Volynkin; 39, Macohasa tetraspila, paratype, Thailand, Chantaburi prov., slide AV4186m Volynkin; 40, Macohasa dimorpha, Philippines, PalaWan, slide MWM 31778 Volynkin; 41, Macotasa biplagella, slide BMNH(E) Arct-2849 (©NHMUK).