Lupinus luisanae var. ocetensis Contreras-Ortiz & Jara, 2018

Contreras-Ortiz, N., Jara-Muñoz, O. A. & Hughes, C. E., 2018, The acaulescent rosette species of Lupinus L. (Fabaceae) of Colombia and Ecuador including a new species from Colombia, Phytotaxa 364 (1), pp. 61-70 : 66-67

Lupinus luisanae var. ocetensis Contreras-Ortiz & Jara , var. nov. Fig. 2 (D–F)

Type:— COLOMBIA. Boyacá: Mpio. Monguí, Páramo de Ocetá, 5°42′22″N, 72°48′45″W, 3700 m, 2 Dec 2014, N. Contreras-Ortiz, G. Ortiz & N. Yepes 50 (holotype ANDES!, isotype Z!).

Lupinus luisanae var. ocetensis differs from Lupinus luisanae var. luisanae by its significantly larger plant size, very narrow elliptic and usually longer leaflets (up to 26.8 cm vs. up to 12 cm long), triangular flexuose free stipule tips (as opposed to lanceolate and flexuose), the standard petal with a white blotch restricted only to the central tip of the petal (compared to a visible white vertical blotch comprising almost one third of the petal width).

Acaulescent rosettes of ca. 80 cm in diameter and up to 1.3 m tall including the inflorescence at full development, with prostrate stolons and secondary branches forming independent secondary plants surrounding the main stem. Stems, scapes and calyces covered by a sparse appressed silky indumentum of 2.8–5 mm long hairs and a dense layer of hoary trichomes 0.3–1 mm long. Free stipule tips triangular, flexuose, (6–) 7–20 × 2–5.1 mm; petioles and stipules covered by a silky indumentum of 6.1–7.9 mm long hairs and a dense layer of hoary trichomes ca. 0.2–0.3 mm long; leaflets very narrowly elliptic, up to 21.4 (–26.8) × 2.7 (–4.8) cm, with connections between secondary veins evident towards the leaflet margin. Inflorescences with ca. 40–45 crowded whorls of 12–14 flowers per whorl; pedicels 5.7–8.7 mm long, covered by an erect silky pubescence of 2.8–5 mm long hairs and a dense layer of hoary trichomes 0.3–1 mm long; corolla with the standard petal with an incipient white central blotch restricted to the tip of the petal. Pods with a yellow stigma persistent at fruit maturity.

Distribution and habitat: — Lupinus luisanae var. ocetensis is endemic to the Páramo of Ocetá, municipality of Monguí, department of Boyacá, Colombia ( Fig. 1 ). This is a grass páramo between 3500 and 3800 m, where Espeletia incana Cuatrecasas (1940b: 435) and Espeletia boyacensis Cuatrecasas (1973: 176) are common.

Etymology: —The infraspecific epithet refers to the name of the Páramo where the taxon is found.

Paratypes:― COLOMBIA. Boyacá: Mpio. de Monguí, Páramo de Ocetá , 5°41′38.4″N, 72°49′31.4″W, 3600 m, 3 Mar 2013, N. Contreras-Ortiz, S. Madriñán, M. P. Contreras, N. Nuerk 20 ( ANDES) ; 5°42′22″N, 72°48′45″W, 3700 m, 2 Dec 2014, N. Contreras-Ortiz, G. Ortiz & N. Yepes 49 ( ANDES, Z) ; 5°42′22″N, 72°48′45″W, 3700 m, 2 Dec 2014, N. Contreras-Ortiz, G. Ortiz & N. Yepes 51 ( ANDES, Z) .

Figures

FIGURE 1. Map of the distributionof the acaulescentrosette species ofLupinus fromColombia and Ecuador.Data pointsare georeferenced herbarium specimen records.

FIGURE 2. Lupinus luisanae Contreras-Ortiz & Jara var. luisanae. A. Detail of inflorescence. B. Habit, Páramo de Pisba. C. Habit, Sierra Nevada del Cocuy. Lupinus luisanae var. ocentesis Contreras-Ortiz & Jara. D. Detail of inflorescence. E. Habit, Páramo de Ocetá.F. Detail of leaf, scale bar 3 cm. Photographs A and B, Astrid Caro; C–F, Natalia Contreras.