Lispe glauca, Pont, 2019

Pont, Adrian C., 2019, Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797, Zootaxa 4557 (1), pp. 1-232 : 139-144

Lispe glauca sp. nov.

( Figs 295–303 )

Diagnosis. In the group of species with a seta on posterior apical margin of hind coxa, L. glauca is most similar to Lispe floccosa but can be distinguished by the relatively simple hind tarsomeres ( Fig. 297 ).

Etymology. The species name is the feminine form Latin adjective “glaucus” (= grey”) and refers to the general appearance of the species.

Material examined. Holotype ♂. AUSTRALIA: Western Australia: Thomas River estuary, Esperance District , Malaise trap, 8.xi.1977, D.H. Colless ( ANIC) . Paratype 1♂. AUSTRALIA: Western Australia: Cranmore Park , bred, xi.1933, M.E. Fuller , 1♂ (very damaged) ( ANIC) .

Description ♂ (♀ not known). Head ( Fig. 296 ). Ground-colour black. Frons at broadest point 0.4, at lunula 0.35, of maximum head-width (frontal view). Fronto-orbital plate uniformly pruinose, together with frontal

triangle, parafacial, face, gena and most of occiput silvery-white pruinose; frontal triangle very broad, frontal vitta totally suppressed. Lower orbital seta short. Parafacial broad, as broad as postpedicel; bare on upper third. Antenna black, only apical edge of pedicel obscurely orange; postpedicel short, 2.0 times as long as broad, falling short of mouth-margin by its length. Arista plumose, the hairing at widest point equal to width of postpedicel. Vibrissa strong. Gena broad, 0.26 of vertical eye-length; genal setae sparse anteriorly and dense behind. Palpus brown, stalk yellow, strongly swollen at tip (as in L. caespitosa , Fig. 286 ).

Thorax. Ground-colour black. Scutum and scutellum rather unevenly light grey dusted; postpronotal lobe, notopleuron and pleura thinly grey dusted. Viewed from in front, scutum wholly silvery dusted; in posterior view, with the grey dust very restricted, limited to a light grey band along suture, running from notopleuron to dc, and with a pair of post grey vittae between dc and ia that coalesce posteriorly to form a broad grey median patch from 2nd post dc to scutellum; scutellum grey dusted. Anterior spiracle yellowish. Acr in 5 irregular rows at suture. Dc 2+3. 2 pprn. 1+2 ia. 1+1 sa, both strong. 1 strong and 1 weaker proepisternal setae. 1 strong proepimeral, slightly shorter than the strong proepisternal, with 9–11 adjacent setulae. Anepisternum with a very short seta in upper anterior corner; posterior row with 3 strong and 2 weak setae. 3 strong kepst, lower one closer to posterior than to anterior. Meron bare below spiracle and above hind coxa. Scutellum with disc densely setulose, 2 irregular rows descending on to sides, otherwise sides and ventral surface bare.

Legs. Black, knees hardly reddish. Fore coxa bare behind. Fore femur with only short setulae on av surface; pv row complete, the setae long and fine, longer than femoral depth, interspersed with multiserial v to pv setulae; pd row complete. Fore tibia without submedian seta; with d, pd and pv apical setae, and also with a distinct ad apical. Fore tarsomeres 2–5 rather short, none with any modifications. Mid femur with very short av setae in basal half, continued as a row of spinulose setulae; pv surface with longer setae in basal half, as long as femoral depth, continuing as a row of erect fine setulae; basal half also with a row of short strong a setae, culminating in a strong seta at middle; 2 p preapicals. Mid tibia with 1 strong p seta; with strong ad, av, v, pv and p apicals, the p short, the ad apical a little before apex. Mid tarsomeres without modifications. Hind coxa with a seta on posterior apical margin. Hind femur with 5 av setae in apical half, the longest longer than femoral depth, preceded by several erect setulae; pv surface with several short setae in basal half, merging into a row of dense fine setulae; ad row complete; 1 d preapical. Hind tibia with 1 ad, with the ground-setulae on this surface rather long; 1 av seta (with an adventitious short second seta on one side); ad and d preapical setae long and strong; 1 av and 1 pv apical setae. Hind tarsus with basal tarsomere not swollen, with a uniform covering of short dense brush-like setulae along p to pv surfaces ( Fig. 297 ).

Wing. Clear. Tegula black, basicosta yellow. Crossvein r-m placed slightly apicad of the point where vein R1 enters costa; dm-cu straight, longer than apical section of vein CuA1. Vein M running straight to wing-margin. Calypters white, margins creamy. Knob of haltere black.

Abdomen. Ground-colour black. Tergites grey dusted, slightly damp in the holotype but apparently unmarked except for very weakly indicated dull spots on tergites 3 and 4 ( Fig 298 ); the paratype with syntergite 1+2 wholly dark. Epandrium grey dusted. Sternites 1–4 grey dusted, sternite 5 shining. Setae only present on tergite 5, with 1 pair of median and 3 of lateral discals, and 5 pairs of erect marginals. Sternites 2–4 with sparse setulae ( Fig. 299 ).

Ƌ terminalia. Unusually large (cercal plate drawn at the same scale as the other species, but lateral view and aedeagus at a smaller scale). Epandrium separated from tergite 5 by syntergosternite 8, which has 2 spiracles; without large setae; weakly produced lobe-like ventrally. Tergite 6 absent. Sternite 6 present as 2 small symmetrical plates, withdrawn beneath sternite 5, and connected to sternite 5 by a pair of broad projections from the sternite. Surstylus absent, i.e. fused to epandrium without trace ( Fig. 300 ). Cercal plate divided dorsally and ventrally ( Fig. 301 ). Hypandrium attached at two points: to a rod originating from wall of epandrium close to upper outer edge of cercal plate; and via a tiny apparent praegonite to lower anterior corner of epandrium. Phallic complex ( Fig 302– 303 ): hypandrium with a pair of long finger-like processes at posterior end that meet and unite with base of epiphallus; praegonite and postgonite present, the latter very long and incurved with their tips crossing ( Fig. 303 ); phallapodeme large, at mid-length joined to hypandrium by a small bridge; phallus long, tube-like, simple, juxta large, without spinules, with a pair of small flap-like processes flanking at base.

Measurements. Wing-length, 5.5 mm. Body-length, 7.0 mm.

Biology. Not known.

Distribution. Known only from Australia (WA).

Figures

FIGURE 295. Lispe glauca sp. nov., ♂ habitus (holotype).

FIGURES 296–299. Lispe glauca sp. nov., ♂. 296. Head, lateral view. 297. Tip of hind tibia and hind tarsomere 1, posterior view. 298. Abdomen, dorsal view. 299. Sternites 2–5. (Holotype.)

FIGURES 300–301. Lispe glauca sp. nov., ♂. 300. Hypopygium, lateral view. 301. Cercal plate, dorsal view. (Holotype.)

FIGURES 302–303. Lispe glauca sp. nov., ♂. 302. Phallic complex, lateral view. 303. Postgonites ventral view. phphallapodeme. (Holotype.)

FIGURES 285–289. Lispe caespitosa sp. nov., ♂. 285. Head, lateral view. 286. Palpus. 287. Tip of hind tibia and hind tarsomere 1, anterior view. 288. Abdomen, dorsal view. 289. Abdominal tergites 4 and 5, lateral view. (285, paratype from V, Pink Lakes; 286–289, paratype from V, Lake Tyrrell.)