Lesticus laevis Fedorenko, 2022

Fedorenko, D. N., 2022, New or little-known Lesticus Dejean, 1828 (Coleoptera: Carabidae: Pterostichini) from Vietnam, Russian Entomological Journal 31 (3), pp. 260-275 : 272-274

9. Lesticus laevis Fedorenko , sp.n.

Figs 12 , 24 , 44–45 .

MATERIAL. Holotype ♂ and paratype ♀ ( ZIN), with handwritten label ‘N. Vietnam / Tamdao [= Tam Dao], 18 V.1995 Gorochov [= Gorokhov]’. Paratype ♂ ( MPSU), wrongly labelled ‘Вьетнам [= Vietnam] / округ Бао- Лок [= Bao Loc District] / 5–23.XI.1992, leg. СинЯев [= Sinyaev]’. — Internal sac of aedeagus examined in the two males .

DIAGNOSIS. A slender, large-sized, apterous species with short metepisterna. At least forebody dorsum bright metallic. Protarsomere 1 with a rudimentary ventral pad in female, metatarsus with a sharp anterolateral carina. Internal sac of aedeagus very similar to that of L. lakhonus ( Figs 44– 45 ).

REDESCRIPTION. Slightly different from L. lakhonus in the following characters: Body large, BL 26.2–28 mm. Elytra shiny, purple green, with reflexed lateral margin violaceous, to nearly black in male, black and dull in apical two fifths and along sides, otherwise shiny, in female. Microsculpture imperceptible on head and pronotum, or very superficial yet traceable on pronotum in female. Dorsal micropunctation almost indistinct.

Head ( Fig. 12 ). Frontal sulci long, deep before the level of anterior supra-ocular seta, very shallow behind. Antennae slightly longer, with about three apical segments surpassing elytral base. Maxillary and labial palps slender: terminal maxillary palpomere parallel-sided or almost so, 0.27 (♀) or 0.30 (♂) times as wide as long; terminal labial palpomere subtriangular, 0.40 (♀) or 0.54–0.59 (♂) times as wide at apex as long.

Pronotum ( Fig. 24 ) almost flat at base so that basal foveae are very shallow, flat, merging into or barely separated from basal dilation of lateral bead, with a few imperceptible punctures only. Inner basal sulci very shallow except at the very base.

Elytra longer in male than in female (Tab. 1), sides slightly more rounded than in L. lakhonus . Basal ridge inwardly reaching stria 3. Interval 3 with 3–4 discal setae, in the latter case apical three situated in stria 2. USS: 24–27.

Table 1. Body ratios in species of Lesticus . Таблица 1. Индексы пропорций тела видов Lesticus .

Metepisternum short, est3L/W 1.0–1.06.

Legs. Meso- and metatarsi with sharp lateral carina. Mesofemur with 3–4 anteroventral setae.

GEOGRAPHIC DISTRIBUTION. Only known from the type locality, Tam Dao , Vinh Phuc Province, Northern Vietnam. Male paratype from ‘Bao Loc District’ is most likely to originate from northern Vietnam, too .

NAME. Refers to smooth basal foveae of the pronotum.

HABITATS AND HABITS. No data.

COMMENTS. The new species is very similar to L. lakhonus , which suggests that it is a montane derivative of that species.

Acknowledgements. I am very grateful to Dr. Boris Kataev (ZISP) and Dr. Kirill Makarov (MPSU) for the loan of material under their care. This study was funded by the Presidium of the Russian academy of sciences, Program No.41 “Biodiversity of natural systems and biological resources of Russia ”.

Figures

Figs 1–12. Head: 1–2 — Lesticus nubilus, 2♂♂; 3 — L. attenuatus sp.n., ♂ paratype; 4 — L. p. praestans, ♂ from Cuc Phuong NP, Vietnam; 5 — L. p. mouhotii, ♀ from Mondulkiri Prov., Cambodia; 6 — L. auricollis, ♂ from Bac Giang Prov., Vietnam; 7–8 — L. verticillatus sp.n., ♂ paratype from Song Thanh NP and ♂ holotype; 9 — L. bellus sp.n., ♀ holotype; 10 — L. buqueti, ♀; 11 — L. lakhonus, ♀; 12 — L. laevis sp.n., ♂ holotype; 1, 10–11 — Cat Tien NP, Vietnam; 2 — Mae Vong env., Thailand. Рис. 1–12. Голова: 1–2 — Lesticus nubilus, 2♂♂; 3 — L. attenuatus sp.n., паратип ♂; 4 — L. p. praestans, ♂ иЗ Нац. парка Кыкфыонг, Вьетнам; 5 — L. p. mouhotii, ♀ иЗ пров. Мондулкири, Камбоджа; 6 — L. auricollis, ♂ иЗ пров. БакЗЯн, Вьетнам; 7–8 — L. verticillatus sp.n., паратип ♂ иЗ нац. парка Сонгтхань и голотип ♂; 9 — L. bellus sp.n., голотип ♀; 10 — L. buqueti, ♀; 11 — L. lakhonus, ♀; 12 — L. laevis sp.n., ♂ голотип; 1, 10–11 — Нац. парк Катьен, Вьетнам; 2 — окр. Мае Вонг, Таиланд.

Figs 13–24. Pronotum: 13–14 — Lesticus nubilus, 2♂♂; 15 — L. attenuatus sp.n., ♂ paratype; 16 — L. p. praestans, ♂ from Cuc Phuong NP, Vietnam; 17 — L. p. mouhotii, ♀ from Mondulkiri Prov., Cambodia; 18 — L. auricollis, ♂ from Bac Giang Prov., Vietnam; 19– 20 — L. verticillatus sp.n., ♂ paratype from Song Thanh NP and ♂ holotype; 21 — L. bellus sp.n., ♀ holotype; 22 — L. buqueti, ♀; 23 — L. lakhonus, ♀; 24 — L. laevis sp.n., ♂ holotype; 13, 22–23 — Cat Tien NP, Vietnam; 14 — Mae Vong env., Thailand. Рис. 13–24. Переднеспинка: 13–14 — Lesticus nubilus, 2♂♂; 15 — L. attenuatus sp.n., паратип ♂; 16 — L. p. praestans, ♂ иЗ Нац. парка Кыкфыонг, Вьетнам; 17 — L. p. mouhotii, ♀ иЗ пров. Мондулкири, Камбоджа; 18 — L. auricollis, ♂ иЗ пров. БакЗЯн, Вьетнам; 19–20 — L. verticillatus sp.n., паратип ♂ иЗ нац. парка Сонгтхань и голотип ♂; 21 — L. bellus sp.n., голотип ♀; 22 — L. buqueti, ♀; 23 — L. lakhonus, ♀; 24 — L. laevis sp.n., ♂ голотип; 13, 22–23 — Нац. парк Катьен, Вьетнам; 14 — окр. Мае Вонг, Таиланд.

Figs 40–45. Aedeagus, median lobe with everted and inflated internal sac: 40–41 — Lesticus buqueti, ♂♂ from Java (40) or Vietnam (41); 42–43 — L. lakhonus; 44–45 — L. laevis sp.n., holotype; 40–41, 43, 45 — right dorsolateral aspect; 42, 44 — left ventrolateral aspect; b, c, d, e — bulbs of internal sac. Scale bars 1 mm. Рис. 40–45. Эдеагус, среднЯЯ долЯ с вывернутым и надутым внутренним меШком: 40–41 — Lesticus buqueti, ♂♂ с Явы (40) и иЗ Вьетнама (41); 42–43 — L. lakhonus; 44–45 — L. laevis sp.n., голотип; 40–41, 43, 45 — справа и сверху; 42, 44 — слева и сниЗу; b, c, d, e — пуЗыри внутреннего меШка. МасШтаб: 1 мм.